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Equity Incentive Plans (Tables) | 9 Months Ended
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Sep. 30, 2020
Share-based Payment Arrangement [Abstract] |
Summary of Stock Option Activity | The following table summarizes stock option activity during the nine months ended September 30, 2020:
| Shares Subject to Options Outstanding |  | Weighted- Average |  |
| Shares |  | Weighted- Average Exercise Price |  | Remaining Contractual Life (Years) |  | Aggregate Intrinsic Value
|  |  |  |  |  |  | (in thousands)
Outstanding as of December 31, 2019 | 8,538,060 |  |  | $ | 13.67 |  |  | 9.0 |  | $ | 35,385 |
Options granted | 2,540,122 |  |  | $ | 14.02 |  |  |  |  |
Option exercised | (76,275) |  |  | $ | 3.73 |  |  |  |  |
Options forfeited/cancelled | (735,012) |  |  | $ | 13.61 |  |  |  |  |
Outstanding as of September 30, 2020 | 10,266,895 |  |  | $ | 13.83 |  |  | 8.4 |  | $ | 19,221 |
Options vested and exercisable as of September 30, 2020 | 3,274,782 |  |  | $ | 12.57 |  |  | 7.7 |  | $ | 10,026 |
Summary of Restricted Stock Activity | The summary of the Company’s restricted stock activity is as follows:
| Number of Restricted Stock Units Outstanding |  | Weighted- Average Grant Date Fair Value
Nonvested at December 31, 2019 | 4,648,526 |  |  | $ | 3.98 |
Granted | 1,176,340 |  |  | $ | 11.94 |
Vested | (1,213,911) |  |  | $ | 3.71 |
Forfeited | (79,957) |  |  | $ | 12.95 |
Nonvested at September 30, 2020 | 4,530,998 |  |  | $ | 5.96 |
Stock-Based Compensation Expense Reported in Condensed Consolidated Statements of Operations | Stock-based compensation expense has been reported in the Company’s condensed consolidated statements of operations as follows (in thousands):
| Three months ended September 30, |  | Nine months ended September 30,
| 2020 |  | 2019 |  | 2020 |  | 2019
Research and development | $ | 4,726 |  |  | $ | 2,727 |  |  | $ | 14,094 |  |  | $ | 6,483 |
General and administrative | 4,197 |  |  | 3,001 |  |  | 11,973 |  |  | 7,474 |
Total stock-based compensation | $ | 8,923 |  |  | $ | 5,728 |  |  | $ | 26,067 |  |  | $ | 13,957 |

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