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Statements of Comprehensive Loss - USD ($) $ in Thousands | 3 Months Ended | 9 Months Ended
--- | --- | ---
Sep. 30, 2015 | Sep. 30, 2014 | Sep. 30, 2015 | Sep. 30, 2014
Revenue |  |  |  |
Collaborations, licenses and milestones | $ 3,503 | $ 848 | $ 6,008 | $ 3,856
Operating expenses |  |  |  |
Research and development | 10,582 | 4,953 | 23,263 | 13,464
General and administrative | 3,233 | 2,182 | 8,521 | 5,499
Total operating expenses | 13,815 | 7,135 | 31,784 | 18,963
Loss from operations | (10,312) | (6,287) | (25,776) | (15,107)
Other income (expenses) |  |  |  |
Interest income | 676 | 2 | 1,067 | 31
Interest expense | (4) | (2) | (11) | (7)
Other income (expenses) | (397) | 9 | (629) | 10
Total other income (expense), net | 275 | 9 | 427 | 34
Net loss | (10,037) | (6,278) | (25,349) | (15,073)
Other comprehensive loss |  |  |  |
Net unrealized gain (loss) on marketable securities available-for-sale | 84 |  | (6) |
Comprehensive loss | $ (9,953) | $ (6,278) | $ (25,355) | $ (15,073)
Basic and diluted net loss per common share (in dollars per share) | $ (0.25) | $ (0.20) | $ (0.66) | $ (0.48)
Weighted-average common shares outstanding used in computing basic and diluted net loss (in shares) | 40,473,520 | 31,395,626 | 38,514,179 | 31,376,502

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