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Statements of Comprehensive Income (Loss) - USD ($)
$ in Thousands
 12 Months Ended
Dec. 31, 2015
Dec. 31, 2014
Dec. 31, 2013
Revenue      
Collaborations, licenses and milestones $ 27,762 $ 9,520 $ 10,172
Operating expenses      
Research and development 34,140 18,516 17,000
General and administrative 11,960 7,461 3,692
Total operating expenses 46,100 25,977 20,692
Loss from operations (18,338) (16,457) (10,520)
Other income (expenses)      
Interest income 1,840 33 14
Interest expense (13) (9) (1,213)
Other income (expense) (1,081) 11 16
Loss on settlement of convertible promissory notes     (48,556)
Total other income (expenses), net 746 35 (49,739)
Net loss (17,592) (16,422) (60,259)
Net deemed contribution on exchange of preferred stock     144,765
Net income (loss) attributable to common stockholders (17,592) (16,422) 84,506
Other comprehensive loss      
Net unrealized loss on marketable securities available-for-sale (516)    
Comprehensive loss $ (18,108) $ (16,422) $ (60,259)
Net income (loss) per share attributable to common shareholders:      
Basic (in dollars per share) $ (0.45) $ (0.52) $ 34.18
Diluted (in dollars per share) $ (0.45) $ (0.52) $ (3.85)
Weighted average shares used to compute net income (loss) per share attributable to common stockholders:      
Basic (in shares) 39,015,131 31,390,631 2,472,581
Diluted (in shares) 39,015,131 31,390,631 15,645,789