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Statements of Comprehensive Income (Loss) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2017 | Dec. 31, 2016 | Dec. 31, 2015
Revenue |  |  |
Collaborations, licenses and milestones | $ 35,711 | $ 87,520 | $ 27,762
Operating expenses |  |  |
Research and development | 71,772 | 51,872 | 34,140
General and administrative | 17,501 | 13,108 | 11,960
Total operating expenses | 89,273 | 64,980 | 46,100
Income (loss) from operations | (53,562) | 22,540 | (18,338)
Other income (expense) |  |  |
Interest income | 4,194 | 2,091 | 744
Interest expense | (13) | (21) | (13)
Other income | (7) | 6 | 15
Total other income, net | 4,174 | 2,076 | 746
Income (loss) before income taxes | (49,388) | 24,616 | (17,592)
Income tax expense (benefit) | (463) | 991 |
Net income (loss) | (48,925) | 23,625 | (17,592)
Other comprehensive income (loss) |  |  |
Net unrealized loss on marketable securities available-for-sale | (367) | (925) | (516)
Comprehensive income (loss) | $ (49,292) | $ 22,700 | $ (18,108)
Net income (loss) per share attributable to common shareholders: |  |  |
Basic net income (loss) (in dollars per share) | $ (1.05) | $ 0.57 | $ (0.45)
Diluted net income (loss) (in dollars per share) | $ (1.05) | $ 0.56 | $ (0.45)
Weighted average shares used to compute net income (loss) per share attributable to common stockholders: |  |  |
Basic (in shares) | 46,817,756 | 41,267,329 | 39,015,131
Diluted (in shares) | 46,817,756 | 42,388,867 | 39,015,131

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