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Stock-Based Compensation - Employee expense (Details) - USD ($) $ in Thousands | 3 Months Ended | 9 Months Ended
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Sep. 30, 2018 | Sep. 30, 2017 | Sep. 30, 2018 | Sep. 30, 2017
Stock-based compensation |  |  |  |
Total employee, director and non-employee stock-based compensation expense | $ 6,120 | $ 3,605 | $ 15,473 | $ 10,211
Employee stock options |  |  |  |
Stock-based compensation |  |  |  |
Total employee, director and non-employee stock-based compensation expense | 5,832 | 3,462 | 14,723 | 9,820
ESPP |  |  |  |
Stock-based compensation |  |  |  |
Total employee, director and non-employee stock-based compensation expense | 209 | 143 | 562 | 391
Restricted stock units |  |  |  |
Stock-based compensation |  |  |  |
Total employee, director and non-employee stock-based compensation expense | 79 |  | 188 |
General and administrative |  |  |  |
Stock-based compensation |  |  |  |
Total employee, director and non-employee stock-based compensation expense | 2,732 | 1,422 | 6,037 | 4,305
Research and development |  |  |  |
Stock-based compensation |  |  |  |
Total employee, director and non-employee stock-based compensation expense | $ 3,388 | $ 2,183 | $ 9,436 | $ 5,906

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- Definition Represents the expense recognized during the period arising from equity-based compensation arrangements (for example, shares of stock, unit, stock options or other equity instruments) with employees, directors and certain consultants qualifying for treatment as employees. + References Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 718 -SubTopic 10 -Section 50 -Paragraph 1 -Subparagraph (b) -URI http://asc.fasb.org/extlink&oid=109197908&loc=d3e5047-113901 Reference 2: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 718 -SubTopic 10 -Section S99 -Paragraph 1 -Subparagraph (SAB Topic 14.F) -URI http://asc.fasb.org/extlink&oid=115993241&loc=d3e301413-122809 Reference 3: http://fasb.org/us-gaap/role/ref/legacyRef -Publisher FASB -Name Accounting Standards Codification -Topic 718 -SubTopic 10 -Section 50 -Paragraph 2 -Subparagraph (h)(1)(i) -URI http://asc.fasb.org/extlink&oid=109197908&loc=d3e5070-113901 + Details
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- Definition Line items represent financial concepts included in a table. These concepts are used to disclose reportable information associated with domain members defined in one or many axes to the table. + References No definition available. + Details
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