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Stock-Based Compensation - Employee expense (Details) - USD ($) $ in Thousands	3 Months Ended		9 Months Ended
	Sep. 30, 2018	Sep. 30, 2017	Sep. 30, 2018	Sep. 30, 2017
Stock-based compensation
Total employee, director and non-employee stock-based compensation expense	$ 6,120	$ 3,605	$ 15,473	$ 10,211
Employee stock options
Stock-based compensation
Total employee, director and non-employee stock-based compensation expense	5,832	3,462	14,723	9,820
ESPP
Stock-based compensation
Total employee, director and non-employee stock-based compensation expense	209	143	562	391
Restricted stock units
Stock-based compensation
Total employee, director and non-employee stock-based compensation expense	79		188
General and administrative
Stock-based compensation
Total employee, director and non-employee stock-based compensation expense	2,732	1,422	6,037	4,305
Research and development
Stock-based compensation
Total employee, director and non-employee stock-based compensation expense	$ 3,388	$ 2,183	$ 9,436	$ 5,906

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- Definition

Represents the expense recognized during the period arising from equity-based compensation arrangements (for example, shares of stock, unit, stock options or other equity instruments) with employees, directors and certain consultants qualifying for treatment as employees.

+ References

Reference 1: http://fasb.org/us-gaap/role/ref/legacyRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 718
-SubTopic 10
-Section 50
-Paragraph 1
-Subparagraph (b)
-URI http://asc.fasb.org/extlink&oid=109197908&loc=d3e5047-113901

Reference 2: http://fasb.org/us-gaap/role/ref/legacyRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 718
-SubTopic 10
-Section S99
-Paragraph 1
-Subparagraph (SAB Topic 14.F)
-URI http://asc.fasb.org/extlink&oid=115993241&loc=d3e301413-122809

Reference 3: http://fasb.org/us-gaap/role/ref/legacyRef
-Publisher FASB
-Name Accounting Standards Codification
-Topic 718
-SubTopic 10
-Section 50
-Paragraph 2
-Subparagraph (h)(1)(i)
-URI http://asc.fasb.org/extlink&oid=109197908&loc=d3e5070-113901

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+ References

No definition available.

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