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Statements of Comprehensive Income (Loss) - USD ($)
$ in Thousands
 3 Months Ended 9 Months Ended
Sep. 30, 2018
Sep. 30, 2017
Sep. 30, 2018
Sep. 30, 2017
Revenue        
Collaborations, licenses and milestones $ 29,039   $ 29,039 $ 16,000
Operating expenses        
Research and development 20,953 $ 19,408 70,371 51,376
General and administrative 7,435 4,172 16,955 13,074
Total operating expenses 28,388 23,580 87,326 64,450
Income (loss) from operations 651 (23,580) (58,287) (48,450)
Other income (expenses)        
Interest income, net 2,642 1,045 6,279 3,134
Other income (expense) (143) 56 (202) 86
Total other income, net 2,499 1,101 6,077 3,220
Net income (loss) before income taxes 3,150 (22,479) (52,210) (45,230)
Income tax expense   173   623
Net income (loss) 3,150 (22,652) (52,210) (45,853)
Other comprehensive income (loss)        
Net unrealized gain (loss) on marketable securities (330) 143 (530) 344
Comprehensive income (loss) $ 2,820 $ (22,509) $ (52,740) $ (45,509)
Net income (loss) per share attributable to common shareholders:        
Basic net income (loss) (in dollars per share) $ 0.06 $ (0.48) $ (0.98) $ (0.98)
Diluted net income (loss) (in dollars per share) $ 0.05 $ (0.48) $ (0.98) $ (0.98)
Weighted average shares used to compute net income (loss) per share attributable to common stockholders:        
Basic (in shares) 55,974,080 46,929,498 53,165,774 46,766,562
Diluted (in shares) 58,313,002 46,929,498 53,165,774 46,766,562