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Prior-Period Financial Statements - Comprehensive Income (Loss) (Details) - USD ($)
$ / shares in Units, $ in Thousands
 3 Months Ended 9 Months Ended 12 Months Ended
Sep. 30, 2018
Sep. 30, 2017
Sep. 30, 2018
Sep. 30, 2017
Dec. 31, 2017
Revenue          
Collaborations, licenses and milestones $ 29,039   $ 29,039 $ 16,000  
Operating expenses          
Research and development 20,953 $ 19,408 70,371 51,376  
General and administrative 7,435 4,172 16,955 13,074  
Total operating expenses 28,388 23,580 87,326 64,450  
Income (loss) from operations 651 (23,580) (58,287) (48,450)  
Other income (expenses)          
Interest income   1,048   3,142  
Interest expense   (3)   (8)  
Other income (expense) (143) 56 (202) 86  
Total other income, net 2,499 1,101 6,077 3,220  
Net income (loss) before income taxes 3,150 (22,479) (52,210) (45,230)  
Income tax expense   173   623  
Net income (loss) 3,150 (22,652) (52,210) (45,853)  
Other comprehensive income          
Net unrealized gain (loss) on marketable securities (330) 143 (530) 344  
Comprehensive income (loss) $ 2,820 $ (22,509) $ (52,740) $ (45,509)  
Basic and diluted net loss (in dollars per share)   $ (0.48)   $ (0.98)  
As Reported Before ASC Topic 606          
Other comprehensive income          
Comprehensive income (loss)         $ (49,292)
ASU 2014-09 | As Reported Before ASC Topic 606          
Revenue          
Collaborations, licenses and milestones   $ 7,090   $ 24,771  
Operating expenses          
Research and development   19,408   51,376  
General and administrative   4,172   13,074  
Total operating expenses   23,580   64,450  
Income (loss) from operations   (16,490)   (39,679)  
Other income (expenses)          
Interest income   1,048   3,142  
Interest expense   (3)   (8)  
Other income (expense)   56   86  
Total other income, net   1,101   3,220  
Net income (loss) before income taxes   (15,389)   (36,459)  
Income tax expense   173   623  
Net income (loss)   (15,562)   (37,082)  
Other comprehensive income          
Net unrealized gain (loss) on marketable securities   143   344  
Comprehensive income (loss)   $ (15,419)   $ (36,738)  
Basic and diluted net loss (in dollars per share)   $ (0.33)   $ (0.79)  
ASU 2014-09 | Effect of Adoption of ASC Topic 606          
Revenue          
Collaborations, licenses and milestones   $ (7,090)   $ (8,771)  
Operating expenses          
Income (loss) from operations   (7,090)   (8,771)  
Other income (expenses)          
Net income (loss) before income taxes   (7,090)   (8,771)  
Net income (loss)   (7,090)   (8,771)  
Other comprehensive income          
Comprehensive income (loss)   $ (7,090)   $ (8,771)  
Basic and diluted net loss (in dollars per share)   $ (0.15)   $ (0.19)