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Prior-Period Financial Statementss - (Details) - USD ($) $ in Thousands | 3 Months Ended | 12 Months Ended
--- | --- | ---
Dec. 31, 2018 | Sep. 30, 2018 | Dec. 31, 2017 | Jun. 30, 2017 | Mar. 31, 2017 | Jun. 30, 2016 | Dec. 31, 2018 | Dec. 31, 2017 | Dec. 31, 2016
Revenue |  |  |  |  |  |  |  |  |
Collaborations, licenses and milestones | $ 11,564 | $ 29,039 | $ 30,150 | $ 12,500 | $ 3,500 |  | $ 40,603 | $ 46,150 | $ 109,020
Research collaboration |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |  |  |
Collaborations, licenses and milestones |  |  |  |  |  |  | $ 20,100 | 20,100 | 34,200
Licensing |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |  |  |
Collaborations, licenses and milestones |  |  |  |  |  |  |  | 100 | 69,800
ASU 2014-09 | Effect of Adoption of ASC Topic 606 |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |  |  |
Collaborations, licenses and milestones |  |  |  |  |  |  |  | 10,439 | 21,500
ASU 2014-09 | Research collaboration |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |  |  |
Collaborations, licenses and milestones |  |  |  |  |  |  |  | $ 20,500 | $ 6,250
ASU 2014-09 | Licensing |  |  |  |  |  |  |  |  |
Revenue |  |  |  |  |  |  |  |  |
Prior revenue recognition period |  |  |  |  |  | 5 years |  |  |
Collaborations, licenses and milestones |  |  |  |  |  | $ 11,300 |  |  |

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