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Income Taxes - Deferred tax assets and liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2018 | Dec. 31, 2017
--- | --- | ---
Deferred income tax assets |  |
Net operating loss carryforwards | $ 49,889 | $ 25,565
Research credits | 23,151 | 16,642
Depreciation | 207 | 437
Unrealized loss on securities | 269 | 504
Accrued compensation | 1,097 | 748
Deferred revenue | 11,222 | 16,820
State taxes |  | (2)
Gross deferred income tax assets | 85,835 | 60,714
Valuation allowance | (82,537) | (57,663)
Net deferred income tax assets | 3,298 | 3,051
Deferred income tax liabilities |  |
Patent costs | (3,142) | (2,873)
Licensing costs | (125) | (142)
Capitalized legal costs | (31) | (36)
Gross deferred income tax liabilities | $ (3,298) | $ (3,051)

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