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Collaborative and Licensing Agreements - Revenue Recognition (Details) - USD ($) $ in Thousands | 3 Months Ended
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Mar. 31, 2021 | Mar. 31, 2020
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | $ 33,965 | $ 32,385
Deferred revenue | 77,800 | 46,200
Research collaboration |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | 14,800 | 1,000
Milestone |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | 12,500 | 12,500
Licensing |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded |  | 15,600
Royalty |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | 6,700 | 3,300
Aimmune |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded |  | 9,600
Alexion |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | 5,300 | 3,300
Astellas |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded |  | 300
Genentech |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | 200 | 700
Gilead |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded |  | 6,000
Janssen |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | 14,600 |
MorphoSys |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |
Revenue recorded | $ 13,900 | $ 12,500

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