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Stock-Based Compensation - Employee expense (Details) - USD ($) $ in Thousands | 3 Months Ended
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Mar. 31, 2021 | Mar. 31, 2020
Stock-based compensation |  |
Total employee, director and non-employee stock-based compensation expense | $ 8,293 | $ 6,512
Employee stock options |  |
Stock-based compensation |  |
Total employee, director and non-employee stock-based compensation expense | 6,530 | 5,882
ESPP |  |
Stock-based compensation |  |
Total employee, director and non-employee stock-based compensation expense | 248 | 194
Restricted stock units |  |
Stock-based compensation |  |
Total employee, director and non-employee stock-based compensation expense | 1,515 | 436
General and administrative |  |
Stock-based compensation |  |
Total employee, director and non-employee stock-based compensation expense | 2,737 | 2,291
Research and development |  |
Stock-based compensation |  |
Total employee, director and non-employee stock-based compensation expense | $ 5,556 | $ 4,221

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