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Statements of Comprehensive Income (Loss) - USD ($)
$ in Thousands
 3 Months Ended
Mar. 31, 2021
Mar. 31, 2020
Revenue    
Collaborations, licenses, milestones, and royalties $ 33,965 $ 32,385
Operating expenses    
Research and development 41,411 33,943
General and administrative 8,226 7,219
Total operating expenses 49,637 41,162
Loss from operations (15,672) (8,777)
Other income (expense)    
Interest income, net 215 3,039
Other income (expense), net 12,970 (2,336)
Total other income, net 13,185 703
Net loss (2,487) (8,074)
Other comprehensive income (loss)    
Net unrealized gain (loss) on marketable securities available-for-sale 23 (105)
Comprehensive loss $ (2,464) $ (8,179)
Net loss per share attributable to common stockholders:    
Basic and diluted net loss (in dollars per share) $ (0.04) $ (0.14)
Weighted average shares used to compute net loss per share attributable to common stockholders:    
Basic and diluted (in shares) 57,997,313 56,946,714