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Marketable Debt and Equity Securities - Equity Securities with Readily Determinable Fair Value (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
--- | --- | ---
Equity Securities |  |
Equity Securities with Readily Determinable Fair Value | $ 36,860 | $ 5,303
Common Stock |  |
Equity Securities |  |
Equity Securities with Readily Determinable Fair Value | 36,860 | 5,303
Astria/Catabasis | Common Stock |  |
Equity Securities |  |
Equity Securities with Readily Determinable Fair Value | 3,449 |
INmune | Common Stock |  |
Equity Securities |  |
Equity Securities with Readily Determinable Fair Value | 19,233 |
MiRagen/Viridian | Common Stock |  |
Equity Securities |  |
Equity Securities with Readily Determinable Fair Value | $ 14,178 | $ 5,303

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- Definition Amount of investment in equity security measured at fair value with change in fair value recognized in net income (FV-NI), classified as current. + References Reference 1: http://www.xbrl.org/2003/role/disclosureRef -Publisher FASB -Name Accounting Standards Codification -Topic 825 -SubTopic 10 -Section 45 -Paragraph 1A -URI http://asc.fasb.org/extlink&oid=123594786&loc=SL75136599-209740 + Details
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