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Income Taxes - Deferred tax assets and liabilities (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
--- | --- | ---
Deferred income tax assets |  |
Net operating loss carryforwards | $ 46,629 | $ 56,182
Research credits | 48,128 | 38,047
Unrealized loss on securities | 327 | 195
Capitalized lease assets | 489 | 288
Accrued compensation | 9,207 | 8,464
Deferred revenue |  | 11,925
Gross deferred income tax assets | 104,780 | 115,101
Valuation allowance | (93,580) | (105,995)
Net deferred income tax assets | 11,200 | 9,106
Deferred income tax liabilities |  |
Patent costs | (3,416) | (4,219)
Equity investment | (3,508) | (4,497)
Licensing costs | (151) | (194)
Capitalized legal costs | (13) | (21)
Depreciation | (288) | (151)
Unrealized gain on securities | (3,824) | (24)
Gross deferred income tax liabilities | $ (11,200) | $ (9,106)

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