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Collaboration and Licensing Agreements (Tables) | 12 Months Ended
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Dec. 31, 2021
Collaboration and Licensing Agreements |
Schedule of revenue by licensees | The $275.1 million, $122.7 million, and $156.7 million of revenue recorded for the years ended December 31, 2021, 2020, and 2019, respectively, were earned principally from the following licensees (in millions): ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | Year Ended |
​ | ​ | December 31, |
​ | ​ | 2021 | ​ | 2020 | ​ | 2019 |
Aimmune |  | $ | — |  | $ | 9.6 |  | $ | — | ​
Alexion | ​ | ​ | 22.2 | ​ |  | 26.2 | ​ |  | 13.0 | ​
Amgen | ​ | ​ | — | ​ |  | — | ​ |  | 5.0 | ​
Astellas | ​ | ​ | — | ​ |  | 3.5 | ​ |  | 14.0 | ​
Genentech | ​ | ​ | 2.5 | ​ |  | 3.5 | ​ |  | 113.9 | ​
Gilead | ​ | ​ | — | ​ |  | 13.5 | ​ |  | — | ​
Janssen | ​ | ​ | 113.8 | ​ | ​ | — | ​ | ​ | — | ​
MorphoSys | ​ | ​ | 18.4 | ​ |  | 39.0 | ​ |  | — | ​
Novartis | ​ | ​ | 43.1 | ​ |  | — | ​ |  | 10.0 | ​
Omeros | ​ | ​ | — | ​ |  | 5.0 | ​ |  | — | ​
Vir | ​ | ​ | 52.7 | ​ |  | 0.3 | ​ |  | 0.8 | ​
Viridian | ​ | ​ | 7.5 | ​ |  | 6.0 | ​ |  | — | ​
Zenas | ​ | ​ | 14.9 | ​ |  | 16.1 | ​ |  | — | ​
Total | ​ | $ | 275.1 | ​ | $ | 122.7 | ​ | $ | 156.7 | ​
Schedule of disaggregation of revenue | ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | Year Ended
​ | ​ | December 31,
​ | ​ | 2021 | ​ | 2020 | ​ | 2019
Research collaboration |  | $ | 93.0 |  | $ | 4.5 |  | $ | 16.3
Milestone | ​ | ​ | 21.0 | ​ |  | 50.2 | ​ |  | 23.2
Licensing | ​ | ​ | 80.8 | ​ |  | 50.2 | ​ |  | 112.2
Royalties | ​ | ​ | 80.3 | ​ |  | 17.8 | ​ |  | 5.0
Total | ​ | $ | 275.1 | ​ | $ | 122.7 | ​ | $ | 156.7

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- Definition Tabular disclosure of disaggregation of revenue into categories depicting how nature, amount, timing, and uncertainty of revenue and cash flows are affected by economic factor. + References Reference 1: http://www.xbrl.org/2009/role/commonPracticeRef -Publisher FASB -Name Accounting Standards Codification -Topic 606 -SubTopic 10 -Section 50 -Paragraph 5 -URI http://asc.fasb.org/extlink&oid=123351226&loc=SL49130545-203045 + Details
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