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Marketable Debt and Equity Securities - Equity Securities Transactions (Details) $ in Thousands	1 Months Ended							12 Months Ended
	Dec. 31, 2021 USD ($) shares	Sep. 30, 2021 USD ($) shares	Aug. 31, 2021	Jul. 31, 2021 USD ($) shares	Jun. 30, 2021 USD ($) shares	Dec. 31, 2020 USD ($) shares	Oct. 31, 2017 USD ($) shares	Dec. 31, 2021 USD ($) shares	Dec. 31, 2020 USD ($)	Dec. 31, 2017 shares
Equity Securities
Net gains recognized on equity securities | $								$ 39,289	$ 105
Impairment | $								762
Preferred Stock
Equity Securities
Equity Securities without Readily Determinable Fair Value | $	$ 31,262					$ 16,071		$ 31,262	$ 16,071
Astria/Catabasis | Common Stock
Equity Securities
Shares received in noncash transaction					259,206
Shares issued upon conversion					3,580,539
Reverse stock split ratio			0.166
Number of shares								3,839,745
Astria/Catabasis | Preferred Stock
Equity Securities
Shares received in noncash transaction				347	3,928
Number of preferred stock converted (in shares)					3,581
Equity Securities without Readily Determinable Fair Value | $	$ 312			$ 1,100	$ 12,100			$ 312
Impairment | $								$ 800
Technology License Agreement | MiRagen/Viridian
Equity Securities
Shares received in noncash transaction	394,737					322,407
Purchase amount of share options or equity in noncash transaction | $	$ 7,500					$ 6,000
Technology License Agreement | MiRagen/Viridian | Common Stock
Equity Securities
Shares received in noncash transaction	394,737					322,407
Technology License Agreement | Astria/Catabasis | Common Stock
Equity Securities
Reverse stock split ratio			0.166
License Agreement | INmune
Equity Securities
Shares received in noncash transaction							1,585,000
Purchase amount of share options or equity in noncash transaction | $							$ 10,000
Number of shares					192,533
Cash consideration on sale of option | $					$ 15,000
License Agreement | INmune | Common Stock
Equity Securities
Shares received in noncash transaction										1,585,000
Additional equity interests (as a percentage)					10.00%					10.00%
Option to acquire shares							108,000			108,000
Shares issued upon conversion					192,533
Purchase amount of share options or equity in noncash transaction | $		$ 800
Net gains recognized on equity securities | $		$ 900
Number of shares		108,000						1,885,533
Cash consideration on sale of option | $					$ 15,000

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The number of shares converted in a noncash (or part noncash) transaction. Noncash is defined as transactions during a period that do not result in cash receipts or cash payments in the period. "Part noncash" refers to that portion of the transaction not resulting in cash receipts or cash payments in the period.

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