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Statements of Comprehensive Income (Loss) - USD ($)
$ in Thousands
 12 Months Ended
Dec. 31, 2021
Dec. 31, 2020
Dec. 31, 2019
Revenue      
Collaborations, licenses, milestones, and royalties $ 275,111 $ 122,694 $ 156,700
Operating expenses      
Research and development 192,507 169,802 118,590
General and administrative 38,837 29,689 24,286
Total operating expenses 231,344 199,491 142,876
Income (loss) from operations 43,767 (76,797) 13,824
Other income (expenses)      
Interest income, net 849 7,264 13,619
Other income (expense), net (1,274) 95 (256)
Gain on equity securities, net 39,289 105  
Total other income, net 38,864 7,464 13,363
Income (loss) before income tax 82,631 (69,333) 27,187
Income tax expense 0 0 312
Net income (loss) 82,631 (69,333) 26,875
Other comprehensive income (loss)      
Net unrealized gain (loss) on marketable debt securities (1,584) (1,087) 2,132
Comprehensive income (loss) $ 81,047 $ (70,420) $ 29,007
Net income (loss) per share attributable to common stockholders:      
Basic net income (loss) (in dollars per share) $ 1.42 $ (1.21) $ 0.48
Diluted net income (loss) (in dollars per share) $ 1.37 $ (1.21) $ 0.46
Weighted average shares used to compute net income (loss) per share attributable to common stockholders:      
Basic (in shares) 58,379,641 57,212,737 56,531,439
Diluted (in shares) 60,495,455 57,212,737 58,467,880