<SEC-DOCUMENT>0000899243-21-010887.txt : 20210310
<SEC-HEADER>0000899243-21-010887.hdr.sgml : 20210310
<ACCEPTANCE-DATETIME>20210310132315
ACCESSION NUMBER:		0000899243-21-010887
CONFORMED SUBMISSION TYPE:	4
PUBLIC DOCUMENT COUNT:		1
CONFORMED PERIOD OF REPORT:	20210308
FILED AS OF DATE:		20210310
DATE AS OF CHANGE:		20210310

REPORTING-OWNER:	

	OWNER DATA:	
		COMPANY CONFORMED NAME:			Castellano Michael J
		CENTRAL INDEX KEY:			0001326138

	FILING VALUES:
		FORM TYPE:		4
		SEC ACT:		1934 Act
		SEC FILE NUMBER:	811-08081
		FILM NUMBER:		21729025

	MAIL ADDRESS:	
		STREET 1:		C/O LAZARD LLC
		STREET 2:		30 ROCKEFELLER PLAZA
		CITY:			NEW YORK
		STATE:			NY
		ZIP:			10020

ISSUER:		

	COMPANY DATA:	
		COMPANY CONFORMED NAME:			BLACKROCK MUNIHOLDINGS FUND, INC.
		CENTRAL INDEX KEY:			0001034665
		IRS NUMBER:				223508039
		FISCAL YEAR END:			1231

	BUSINESS ADDRESS:	
		STREET 1:		100 BELLEVUE PARKWAY
		CITY:			WILMINGTON
		STATE:			DE
		ZIP:			19809
		BUSINESS PHONE:		800-441-7762

	MAIL ADDRESS:	
		STREET 1:		100 BELLEVUE PARKWAY
		CITY:			WILMINGTON
		STATE:			DE
		ZIP:			19809

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	BLACKROCK MUNIHOLDINGS FUND INC
		DATE OF NAME CHANGE:	20061026

	FORMER COMPANY:	
		FORMER CONFORMED NAME:	MUNIHOLDINGS FUND INC
		DATE OF NAME CHANGE:	19970226
</SEC-HEADER>
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<TYPE>4
<SEQUENCE>1
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<DESCRIPTION>FORM 4 SUBMISSION
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    <periodOfReport>2021-03-08</periodOfReport>

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        <issuerCik>0001034665</issuerCik>
        <issuerName>BLACKROCK MUNIHOLDINGS FUND, INC.</issuerName>
        <issuerTradingSymbol>MHD</issuerTradingSymbol>
    </issuer>

    <reportingOwner>
        <reportingOwnerId>
            <rptOwnerCik>0001326138</rptOwnerCik>
            <rptOwnerName>Castellano Michael J</rptOwnerName>
        </reportingOwnerId>
        <reportingOwnerAddress>
            <rptOwnerStreet1>55 EAST 52ND STREET</rptOwnerStreet1>
            <rptOwnerStreet2></rptOwnerStreet2>
            <rptOwnerCity>NEW YORK</rptOwnerCity>
            <rptOwnerState>NY</rptOwnerState>
            <rptOwnerZipCode>10055</rptOwnerZipCode>
            <rptOwnerStateDescription></rptOwnerStateDescription>
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            <isOfficer>0</isOfficer>
            <isTenPercentOwner>0</isTenPercentOwner>
            <isOther>0</isOther>
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    <footnotes>
        <footnote id="F1">Effective as of March 8, 2021, BlackRock Municipal Income Investment Quality Trust (the &quot;Target Fund&quot;) was reorganized into BlackRock MuniHoldings Fund, Inc. (the &quot;Acquiring Fund&quot;). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $15.2503 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.89746185. In the Reorganization, Michael J. Castellano received 627 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 700 common shares of the Target Fund.</footnote>
        <footnote id="F2">Effective as of March 8, 2021, BlackRock MuniHoldings Fund II, Inc. (the &quot;Target Fund&quot;) was reorganized into BlackRock MuniHoldings Fund, Inc. (the &quot;Acquiring Fund&quot;). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $15.7316 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.92578578. In the Reorganization, Michael J. Castellano received 647 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 700 common shares of the Target Fund.</footnote>
        <footnote id="F3">Effective as of March 8, 2021, BlackRock MuniHoldings Quality Fund, Inc. (the &quot;Target Fund&quot;) was reorganized into BlackRock MuniHoldings Fund, Inc. (the &quot;Acquiring Fund&quot;). In the Reorganization, common shareholders of the Target Fund received common shares of the Acquiring Fund having a value equal to the aggregate NAV of the Target Fund common shares surrendered as determined at the close of business on March 5, 2021, less the costs of the Reorganization. As of March 5, 2021, the Target Fund reported a NAV per share of $13.8681 and the Acquiring Fund reported a NAV per share of $16.9927. The conversion ratio for the Target Fund's common shares was 0.81612104. In the Reorganization, Michael J. Castellano received 652 common shares (and cash for fractional shares, if any) of the Acquiring Fund in exchange for his 800 common shares of the Target Fund.</footnote>
    </footnotes>

    <remarks></remarks>

    <ownerSignature>
        <signatureName>/s/ Dean A. Caruvana as Attorney-in-Fact</signatureName>
        <signatureDate>2021-03-10</signatureDate>
    </ownerSignature>
</ownershipDocument>
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</TEXT>
</DOCUMENT>
</SEC-DOCUMENT>
