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Condensed Consolidated Statements of Shareholders’ Equity $ in Thousands | USD ($) shares | ADDITIONAL PAID-IN CAPITAL USD ($) | ACCUMULATED OTHER COMPREHENSIVE (LOSS)/ INCOME USD ($) | ACCUMULATED DEFICIT USD ($) | Ordinary shares £ / shares shares | Ordinary shares COMMON STOCK USD ($) shares | Deferred shares £ / shares shares | Deferred shares COMMON STOCK USD ($) shares
--- | --- | --- | --- | --- | --- | --- | --- | ---
Beginning balance (in shares) at Dec. 31, 2021 | shares |  |  |  |  |  | 42,019,874,000 |  | 1,000
Beginning balance at Dec. 31, 2021 | $ 284,412 | $ 444,750 | $ 8,840 | $ (169,641) |  | $ 435 |  | $ 28
Exercise of share options (in shares) | shares |  |  |  |  | 376,158 | 376,158,000 |  |
Exercise of share options | 397 | 393 |  |  |  | $ 4 |  |
Vesting of restricted stock units (in shares) | shares |  |  |  |  | 10,659 | 68,534,000 |  |
Vesting of restricted stock units | 1 |  |  |  |  | $ 1 |  |
Share-based compensation expense | 3,128 | 3,128 |  |  |  |  |  |
Unrealized loss on foreign currency translation | (7,193) |  | (7,193) |  |  |  |  |
Net loss | (21,171) |  |  | (21,171) |  |  |  |
Ending balance (in shares) at Mar. 31, 2022 | shares |  |  |  |  |  | 42,464,566,000 |  | 1,000
Ending balance at Mar. 31, 2022 | 259,574 | 448,271 | 1,647 | (190,812) |  | $ 440 |  | $ 28
Common stock, par value (in GBP per share) | £ / shares |  |  |  |  | £ 0.008 |  | £ 21,921.504 |
Beginning balance (in shares) at Dec. 31, 2022 | shares |  |  |  |  | 42,631,794 | 42,631,794,000 | 1 | 1,000
Beginning balance at Dec. 31, 2022 | $ 181,280 | 458,825 | (16,867) | (261,146) |  | $ 440 |  | $ 28
Exercise of share options (in shares) | shares | 3,790 |  |  |  | 3,790 | 3,790,000 |  |
Issuance of ordinary shares under ATM offering, net of issuance costs (in shares) | shares |  |  |  |  |  | 113,420,000 |  |
Issuance of ordinary shares under ATM facility, net of issuance costs | $ 1,188 | 1,187 |  |  |  | $ 1 |  |
Shares tendered for withholding taxes | (109) | (109) |  |  |  |  |  |
Vesting of restricted stock units (in shares) | shares |  |  |  |  | 21,579 | 30,481,000 |  |
Vesting of restricted stock units | 0 | (1) |  |  |  | $ 1 |  |
Share-based compensation expense | 4,071 | 4,071 |  |  |  |  |  |
Unrealized loss on foreign currency translation | (578) |  | (578) |  |  |  |  |
Net loss | (24,208) |  |  | (24,208) |  |  |  |
Ending balance (in shares) at Mar. 31, 2023 | shares |  |  |  |  | 42,779,485 | 42,779,485,000 | 1 | 1,000
Ending balance at Mar. 31, 2023 | $ 161,644 | $ 463,973 | $ (17,445) | $ (285,354) |  | $ 442 |  | $ 28
Common stock, par value (in GBP per share) | £ / shares |  |  |  |  | £ 0.008 |  | £ 21,921.504 |

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