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Stock-Based Compensation - Summary of Stock Option Activity (Details) - 2011 Plan, 2019 Plan and Inducement Plan - USD ($) $ / shares in Units, $ in Thousands | 3 Months Ended | 12 Months Ended
--- | --- | ---
Mar. 31, 2024 | Dec. 31, 2023
Share Based Compensation Arrangement By Share Based Payment Award [Line Items] |  |
Outstanding Options, Number of Shares, Beginning Balance | 5,805,586 |
Outstanding Options, Number of Shares, granted | 2,400,500 |
Outstanding Options, Number of Shares, forfeited or expired | (256,904) |
Outstanding Options, Number of Shares, Ending Balance | 7,949,182 | 5,805,586
Options vested and exercisable, Number of Shares | 3,321,228 |
Outstanding Options, Weighted-Average Exercise Price, Beginning Balance | $ 7.40 |
Outstanding Options, Weighted-Average Exercise Price, granted | 1.59 |
Outstanding Options, Weighted-Average Exercise Price, forfeited or expired | 7.60 |
Outstanding Options, Weighted-Average Exercise Price, Ending Balance | 5.64 | $ 7.40
Options vested and exercisable, Weighted-Average Exercise Price | $ 9.62 |
Outstanding Options, Weighted-Average Remaining Contractual Term (in years) | 7 years 10 months 6 days | 6 years 10 months 24 days
Options vested and exercisable, Weighted-Average Remaining Contractual Term (in years) | 5 years 8 months 19 days |
Outstanding Options, Aggregate Intrinsic Value | $ 46 | $ 64
Performance-Based Stock Option |  |
Share Based Compensation Arrangement By Share Based Payment Award [Line Items] |  |
Outstanding Options, Number of Shares, granted | 264,500 |
Outstanding Options, Number of Shares, Ending Balance | 264,500 |
Outstanding Options, Weighted-Average Exercise Price, granted | $ 1.61 |
Outstanding Options, Weighted-Average Exercise Price, Ending Balance | $ 1.61 |
Outstanding Options, Weighted-Average Remaining Contractual Term (in years) | 9 years 11 months 15 days |

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