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License and Collaboration Agreements (Details) - USD ($) | 12 Months Ended |  |
--- | --- | --- | ---
Dec. 31, 2016 | Dec. 31, 2015 | Feb. 28, 2017 | Feb. 13, 2017
Scripps Research Institute |  |  |  |
License and Collaboration Agreements |  |  |  |
Milestone payments made | $ 0 |  |  |
Vernalis (R&D) Limited | Subsequent Events |  |  |  |
License and Collaboration Agreements |  |  |  |
Milestone payments made |  |  | $ 3,000,000 | $ 0
Licensing Agreement | Scripps Research Institute |  |  |  |
License and Collaboration Agreements |  |  |  |
Minimum annual fee |  | $ 25,000 |  |
Aggregate potential milestone payments | $ 2,600,000 |  |  |
Period of written notice to terminate | 90 days |  |  |
Licensing Agreement | Scripps Research Institute | Research and development |  |  |  |
License and Collaboration Agreements |  |  |  |
One-time cash payment |  | $ 10,000 |  |
Licensing Agreement | Vernalis (R&D) Limited |  |  |  |
License and Collaboration Agreements |  |  |  |
Period of written notice to terminate | 90 days |  |  |
Licensing Agreement | Vernalis (R&D) Limited | Minimum |  |  |  |
License and Collaboration Agreements |  |  |  |
Aggregate potential milestone payments | $ 200,000,000 |  |  |
Licensing Agreement | Vernalis (R&D) Limited | Research and development |  |  |  |
License and Collaboration Agreements |  |  |  |
One-time cash payment | $ 1,000,000 |  |  |

X
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