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License and Collaboration Agreements (Details) - USD ($) | 6 Months Ended |  |  |
--- | --- | --- | --- | ---
Jun. 30, 2020 | Feb. 28, 2017 | Dec. 31, 2015 | Feb. 28, 2015
Monash License |  |  |  |
License and Collaboration Agreements |  |  |  |
Minimum annual fee | $ 25,000 |  |  |
Aggregate potential milestone payments | $ 45,100,000 |  |  |
Period of written notice to terminate | 30 days |  |  |
Monash License | Research and development |  |  |  |
License and Collaboration Agreements |  |  |  |
One-time cash payment | $ 275,000 |  |  |
Licensing Agreement | Scripps Research Institute |  |  |  |
License and Collaboration Agreements |  |  |  |
Minimum annual fee |  |  | $ 25,000 |
Aggregate potential milestone payments | 2,600,000 |  |  |
Milestone payments made | $ 0 |  |  |
Period of written notice to terminate | 90 days |  |  |
Licensing Agreement | Scripps Research Institute | Research and development |  |  |  |
License and Collaboration Agreements |  |  |  |
One-time cash payment |  |  | $ 10,000 |
Licensing Agreement | Vernalis (R&D) Limited |  |  |  |
License and Collaboration Agreements |  |  |  |
Milestone payments made |  | $ 3,000,000.0 |  |
Period of written notice to terminate | 90 days |  |  |
Licensing Agreement | Vernalis (R&D) Limited | Minimum |  |  |  |
License and Collaboration Agreements |  |  |  |
Aggregate potential milestone payments | $ 220,000,000 |  |  |
Licensing Agreement | Vernalis (R&D) Limited | Research and development |  |  |  |
License and Collaboration Agreements |  |  |  |
One-time cash payment |  |  |  | $ 1,000,000.0

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