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Significant agreements - Sanofi Collaboration Agreement (Details) - USD ($) $ in Thousands | 1 Months Ended | 3 Months Ended | 6 Months Ended |  |
--- | --- | --- | --- | --- | ---
Aug. 31, 2017 | Jun. 30, 2020 | Jun. 30, 2019 | Jun. 30, 2020 | Jun. 30, 2019 | Dec. 31, 2019 | Dec. 31, 2018
Significant agreements. |  |  |  |  |  |  |
Revenue recognized by non exercising of option |  | $ 0 | $ 0 |  | $ 5,286 |  |
Revenue from collaborative arrangement |  | 1,571 | 1,522 | $ 2,700 | 7,906 |  |
Deferred revenue |  | 33,082 |  | 33,082 |  | $ 5,657 | $ 14,635
Development milestone |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |
Non refundable payment for services during the BV Bicycle Research Term |  |  |  | 4,200 |  |  |
Sanofi |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |
Total transaction price initially determined | $ 14,200 |  |  |  |  | 14,900 |
Upfront cash payment | 10,000 |  |  |  |  |  |
Non refundable payment for services during the BV Bicycle Research Term | 4,200 |  |  |  |  |  |
Revenue from collaborative arrangement |  | 0 | 57 | 0 | $ 6,016 |  |
Deferred revenue |  | $ 0 |  | $ 0 |  | $ 0 | $ 9,908
Sanofi | Sickle Cell Research License and Related Services |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |
Total transaction price initially determined | 1,405 |  |  |  |  |  |
Sanofi | Hemophilia Research License and Related Services |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |
Total transaction price initially determined | 2,811 |  |  |  |  |  |
Sanofi | Sickle Cell License Option Material Right |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |
Total transaction price initially determined | 5,286 |  |  |  |  |  |
Revenue recognized by non exercising of option |  |  | $ 5,300 |  |  |  |
Sanofi | Hemophilia License Option Material Right |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |
Total transaction price initially determined | $ 4,698 |  |  |  |  |  |

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