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Significant agreements - Oxurion Collaboration Agreement (Details) $ in Thousands, € in Millions | 3 Months Ended | 6 Months Ended |  |  |
--- | --- | --- | --- | --- | ---
Jun. 30, 2020 USD ($) | Jun. 30, 2019 USD ($) | Jun. 30, 2020 EUR (€) | Jun. 30, 2020 USD ($) | Jun. 30, 2019 USD ($) | Jun. 30, 2020 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($)
Significant agreements. |  |  |  |  |  |  |  |
Revenue from collaborative arrangement | $ | $ 1,571 | $ 1,522 |  | $ 2,700 | $ 7,906 |  |  |
Deferred revenue | $ |  |  |  |  |  | $ 33,082 | $ 5,657 | $ 14,635
Oxurion |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Threshold period of notice required for termination of agreement |  |  | 90 days | 90 days |  |  |  |
Revenue from collaborative arrangement | $ | $ 0 | $ 0 |  | $ 0 | $ 0 |  |  |
Deferred revenue | $ |  |  |  |  |  | $ 0 |  |
Oxurion | Development milestone |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Upfront cash payment |  |  | € 1.0 |  |  |  |  |
Potential milestone payments |  |  | 8.3 |  |  |  |  |
Revenue under contracts with customer from milestone payment |  |  | 1.8 |  |  |  |  |
Oxurion | Regulatory milestone |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Potential milestone payments |  |  | 16.5 |  |  |  |  |
Oxurion | Granting first regulatory approval or first indication |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Potential milestone payments |  |  | € 5.0 |  |  |  |  |

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- Definition Amount of obligation to transfer good or service to customer for which consideration has been received or is receivable. + References Reference 1: http://www.xbrl.org/2003/role/disclosureRef -Publisher FASB -Name Accounting Standards Codification -Topic 606 -SubTopic 10 -Section 45 -Paragraph 1 -URI http://asc.fasb.org/extlink&oid=118944033&loc=SL49130531-203044 Reference 2: http://www.xbrl.org/2003/role/disclosureRef -Publisher FASB -Name Accounting Standards Codification -Topic 606 -SubTopic 10 -Section 50 -Paragraph 8 -Subparagraph (a) -URI http://asc.fasb.org/extlink&oid=118956577&loc=SL49130549-203045 Reference 3: http://www.xbrl.org/2003/role/disclosureRef -Publisher FASB -Name Accounting Standards Codification -Topic 606 -SubTopic 10 -Section 45 -Paragraph 2 -URI http://asc.fasb.org/extlink&oid=118944033&loc=SL49130532-203044 + Details
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