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Significant agreements - Oxurion Collaboration Agreement (Details) $ in Thousands, € in Millions | 3 Months Ended | 9 Months Ended |  |  |
--- | --- | --- | --- | --- | ---
Sep. 30, 2020 USD ($) | Sep. 30, 2019 USD ($) | Sep. 30, 2020 EUR (€) | Sep. 30, 2020 USD ($) | Sep. 30, 2019 USD ($) | Sep. 30, 2020 USD ($) | Dec. 31, 2019 USD ($) | Dec. 31, 2018 USD ($)
Significant agreements. |  |  |  |  |  |  |  |
Revenue from collaborative arrangement | $ 3,842 | $ 614 |  | $ 6,542 | $ 8,520 |  |  |
Deferred revenue |  |  |  |  |  | $ 33,275 | $ 5,657 | $ 14,635
Revenue recognized | 1,267 | 173 |  | $ 3,496 | 5,596 |  |  |
Oxurion |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Threshold period of notice required for termination of agreement |  |  | 90 days | 90 days |  |  |  |
Revenue from collaborative arrangement | 2,362 | $ 0 |  | $ 2,362 | $ 0 |  |  |
Deferred revenue |  |  |  |  |  | $ 0 |  |
Revenue recognized | $ 2,400 |  |  | $ 2,400 |  |  |  |
Oxurion | Development milestone |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Upfront cash payment | € |  |  | € 1.0 |  |  |  |  |
Potential milestone payments | € |  |  | 8.3 |  |  |  |  |
Revenue under contracts with customer from milestone payment | € |  |  | 3.8 |  |  |  |  |
Oxurion | Regulatory milestone |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Potential milestone payments | € |  |  | 16.5 |  |  |  |  |
Oxurion | Granting first regulatory approval or first indication |  |  |  |  |  |  |  |
Significant agreements. |  |  |  |  |  |  |  |
Potential milestone payments | € |  |  | € 5.0 |  |  |  |  |

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