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Significant agreements - Summary of Contract Assets and Liabilities (Details) - USD ($) | 12 Months Ended
--- | ---
Dec. 31, 2024 | Dec. 31, 2023
Contract liabilities: |  |
Beginning Balance | $ 135,194,000 | $ 61,873,000
Additions | 3,386,000 | 95,960,000
Deductions | (35,275,000) | (26,976,000)
Impact of Exchange Rates | (1,647,000) | 4,337,000
Ending Balance | 101,658,000 | 135,194,000
Contract assets |  |
Contract assets | 0 | 0
Bayer collaboration deferred revenue |  |
Contract liabilities: |  |
Beginning Balance | 43,618,000 |
Additions | 296,000 | 45,000,000
Deductions | (3,393,000) | (1,160,000)
Impact of Exchange Rates | (561,000) | (222,000)
Ending Balance | 39,960,000 | 43,618,000
Bayer collaboration deferred revenue | Deferred revenue material rights option |  |
Contract liabilities: |  |
Ending Balance | 31,900,000 |
Novartis collaboration deferred revenue |  |
Contract liabilities: |  |
Beginning Balance | 50,008,000 |
Additions | 3,000,000 | 50,000,000
Deductions | (8,161,000) | (1,909,000)
Impact of Exchange Rates | (774,000) | 1,917,000
Ending Balance | 44,073,000 | 50,008,000
Novartis collaboration deferred revenue | Deferred revenue material rights option |  |
Contract liabilities: |  |
Ending Balance | 32,000,000 |
Ionis collaboration deferred revenue |  |
Contract liabilities: |  |
Beginning Balance | 12,464,000 | 21,489,000
Additions | 90,000 | 960,000
Deductions | (8,881,000) | (10,734,000)
Impact of Exchange Rates | (86,000) | 749,000
Ending Balance | 3,587,000 | 12,464,000
Ionis collaboration deferred revenue | Deferred revenue material rights option |  |
Contract liabilities: |  |
Ending Balance | 3,600,000 |
Genentech collaboration deferred revenue |  |
Contract liabilities: |  |
Beginning Balance | 29,104,000 | 39,308,000
Deductions | (14,840,000) | (11,969,000)
Impact of Exchange Rates | (226,000) | 1,765,000
Ending Balance | 14,038,000 | 29,104,000
Genentech collaboration deferred revenue | Deferred revenue material rights option |  |
Contract liabilities: |  |
Ending Balance | $ 13,500,000 |
AstraZeneca collaboration deferred revenue |  |
Contract liabilities: |  |
Beginning Balance |  | 1,076,000
Deductions |  | (1,204,000)
Impact of Exchange Rates |  | $ 128,000

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