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Significant agreements - Ionis Accounting Analysis (Details) $ in Thousands		1 Months Ended			12 Months Ended
	Jul. 09, 2021 USD ($) item	Oct. 31, 2022 USD ($)	Dec. 31, 2021 USD ($)	Jan. 31, 2021 USD ($)	Dec. 31, 2024 USD ($) item	Dec. 31, 2023 USD ($)	Dec. 31, 2022 USD ($)	Sep. 07, 2021 USD ($)	Dec. 31, 2020 USD ($)
Significant agreements.
Collaboration revenue					$ 35,275	$ 26,976	$ 14,463
Deferred revenue					$ 101,658	135,194	61,873
Ionis
Significant agreements.
Number of material rights associated with future payments | item					4
Number of collaboration targets | item	4
Transaction price	$ 38,000				$ 37,800			$ 38,000
Upfront cash payment	31,000
Estimated amount payable to CROs	600				400	600
Collaboration revenue					8,881	10,734	9,347
Deferred revenue					$ 3,587	$ 12,464	$ 21,489
Ionis | Minimum
Significant agreements.
Material rights exercise period					4 years
Ionis | Maximum
Significant agreements.
Material rights exercise period					7 years
Ionis | Combined licenses and research and discovery performance obligation
Significant agreements.
Transaction price								34,100
Ionis | Four material rights associated with credits for IND Acceptance fees
Significant agreements.
Transaction price								$ 3,900
Ionis | Ionis Share Purchase Agreement
Significant agreements.
Transaction price	3,400
Ionis Evaluation and Option Agreement
Significant agreements.
Upfront cash payment	$ 3,000			$ 3,000
Deferred revenue									$ 3,000
Amended Ionis
Significant agreements.
Upfront cash payment			$ 1,600
Collaboration revenue		$ 800	$ 800
Third Amended Ionis
Significant agreements.
Collaboration revenue					$ 800

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