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NIH Contract (Details) - USD ($) $ in Thousands | Mar. 31, 2022 | Dec. 31, 2021
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Research and Development [Abstract] |  |
Cash receipts from milestones achieved | $ 34,016 | $ 34,016
Cumulative amounts applied against operating costs (excluding depreciation) | (4,526) | (4,522)
Cumulative amounts applied against depreciation expense for assets placed in service | (1,510) | (703)
Cumulative amounts recognized as non-operating income | (7,140) | (7,140)
Total deferred grant income | 20,840 | 21,651
Assets placed in service, gross | 20,425 | 16,890
Construction-in-progress | 937 | 3,909
Carrying value of property and equipment, net | 19,852 | 20,096
Estimated future capital expenditures | 988 | 1,555
Deferred grant income, current | 3,603 | 3,535
Deferred grant income, non-current | $ 17,237 | $ 18,116

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