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NIH Contract (Details) - USD ($)
$ in Thousands
Mar. 31, 2022
Dec. 31, 2021
Research and Development [Abstract]    
Cash receipts from milestones achieved $ 34,016 $ 34,016
Cumulative amounts applied against operating costs (excluding depreciation) (4,526) (4,522)
Cumulative amounts applied against depreciation expense for assets placed in service (1,510) (703)
Cumulative amounts recognized as non-operating income (7,140) (7,140)
Total deferred grant income 20,840 21,651
Assets placed in service, gross 20,425 16,890
Construction-in-progress 937 3,909
Carrying value of property and equipment, net 19,852 20,096
Estimated future capital expenditures 988 1,555
Deferred grant income, current 3,603 3,535
Deferred grant income, non-current $ 17,237 $ 18,116