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Related Parties - Additional Information (Detail) - USD ($) $ in Thousands	12 Months Ended
	Dec. 31, 2016	Dec. 31, 2015	Jun. 30, 2012
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Accrued expenses due to related party		$ 1,546
Intrexon [Member]
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Collaboration Agreement cancellation notification	90 days
Collaboration Agreement cancellation with material breach that cannot be remedied	60 days
Accrued expenses due to related party	$ 3,000	1,500
Intrexon [Member] | Commercialization Milestones [Member]
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Commercialization milestones obligation	12,000
Intrexon [Member] | Sales Milestones [Member]
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Sales milestones	$ 22,500
Intrexon [Member] | Research and Development [Member]
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Percentage of reimbursement expenses	50.00%
Percentage of reimbursement expenses subject to acceptance	50.00%
Research and development expense from transaction with Intrexon	$ 3,000	3,100
Purpose, Co. [Member]
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items]
Reimbursement for development cost, amount			$ 300
Reimbursement for development cost, paid	$ 100	$ 200

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- Definition

Amount committed for reimbursement of development costs of new technology.

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- Definition

Carrying amount as of the balance sheet date of obligations due all related parties. For classified balance sheets, represents the current portion of such liabilities (due within one year or within the normal operating cycle if longer).

+ References

Reference 1: http://www.xbrl.org/2003/role/presentationRef
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Expenses recognized resulting from transactions (excluding transactions that are eliminated in consolidated or combined financial statements) with related party.

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