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Share-Based Compensation (Tables)
 3 Months Ended
Mar. 31, 2020
Share-based Payment Arrangement [Abstract]  
Schedule of Stock Options Valuation Assumptions
The assumptions used in the Black-Scholes option pricing model to determine the fair value of the share options granted by the Company during the three months ended March 31, 2020 were as follows:
 
March 31,
 
2020
Expected option life (years)
6.08
Risk-free interest rate
0.58% - 1.66%
Expected volatility
76.38% - 77.64%
Expected dividend yield
—%
Stock Option Rollforward
The table below summarizes activity for the three months ended March 31, 2020:
 
Number of
Options
 
Weighted-Average Exercise Price
 
Weighted-Average Remaining Contractual Term (Years)
 
Aggregate Intrinsic Value
Outstanding as of December 31, 2019
5,936,239

 
$
17.71

 
9.02

 
$
11,873

Granted
96,500

 
$
8.81

 
—

 
—

Exercised
(14,976
)
 
$
1.77

 
—

 
—

Canceled or forfeited
(102,814
)
 
$
24.62

 
—

 
—

Outstanding as of March 31, 2020
5,914,949

 
$
17.49

 
8.79

 
$
2,885

Exercisable as of March 31, 2020
1,735,080

 
$
17.48

 
8.17

 
$
2,013

Vested and expected to vest as of March 31, 2020
5,914,949

 
$
17.49

 
8.79

 
$
2,885
Schedule of Nonvested Share Activity
A summary of the changes in the Company’s restricted ordinary shares during the three months ended March 31, 2020 is as follows:
 
Number of
restricted
shares
 
Weighted average grant date fair value
Unvested and outstanding at December 31, 2019
314,744

 
$
4.22

Granted
—

 
—

Vested
(91,416
)
 
4.32

Canceled or forfeited
(50
)
 
4.44

Unvested and outstanding at March 31, 2020
223,278

 
$
4.18

During the three months ended March 31, 2020, the Company did not grant RSUs under the 2018 Plan. The following is a summary of RSU activity for the 2018 Plan for the three months ended March 31, 2020:
 
Number of
restricted
shares
 
Weighted average grant date fair value
Unvested and outstanding at December 31, 2019
500,000

 
$
12.09

Granted
—

 
—

Vested
—

 
—

Canceled or forfeited
(1,250
)
 
12.09

Unvested and outstanding at March 31, 2020
498,750

 
$
12.09
Schedule Share Based Compensation Expense
Share-based compensation expense recorded as research and development expenses, general and administrative expenses, and capitalized to property and equipment is as follows (in thousands):
 
Three Months Ended March 31,
 
2020
 
2019
Research and development
$
4,562

 
$
4,144

General and administrative
1,627

 
3,221

Capitalized to property and equipment
46

 
—

Total share-based compensation expense
$
6,235

 
$
7,365