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Accrued Expenses and Other Liabilities (Details) $ in Thousands, £ in Millions | Sep. 30, 2020 USD ($) | Dec. 31, 2019 USD ($) | Mar. 31, 2018 GBP (£)
--- | --- | --- | ---
Payables and Accruals [Abstract] |  |  |
Compensation and benefits | $ 5,937 | $ 6,568 |
Research and development costs | 13,073 | 10,449 |
UCLB milestone and option | 840 | 663 | £ 0.5
Professional fees | 5,148 | 2,611 |
U.S. corporate income and local taxes | 0 | 391 |
Other liabilities | 402 | 716 |
Total accrued expenses and other liabilities | $ 25,400 | $ 21,398 |

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