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Accrued Expenses and Other Liabilities (Details) £ in Thousands, $ in Thousands | Sep. 30, 2021 USD ($) | Dec. 31, 2020 USD ($) | Mar. 31, 2018 GBP (£)
--- | --- | --- | ---
Payables and Accruals [Abstract] |  |  |
Compensation and benefits | $ 8,798 | $ 8,732 |
Research and development costs | 16,457 | 15,343 |
UCLB milestone and option | 0 | 205 | £ 350
Professional fees | 3,060 | 3,005 |
U.S. corporate income and local taxes | 7 | 0 |
Other liabilities | 312 | 496 |
Total accrued expenses and other liabilities | $ 28,634 | $ 27,781 |

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