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Income Taxes (Details) - USD ($) $ in Thousands | 12 Months Ended
--- | ---
Dec. 31, 2015 | Dec. 31, 2014 | Dec. 31, 2013
Reconciliation from the U.S. statutory income tax rate to the effective income tax rate |  |  |
Federal income tax (benefit) at statutory rate | 34.00% | 34.00% | 34.00%
Permanent differences | (0.86%) | (1.29%) | (0.50%)
Federal research and development credits and adjustments | 2.64% | 2.70% | 5.80%
State income tax benefit, net of federal benefit | 5.77% | 6.03% | 5.88%
Other (as a percent) | 0.28% | (0.15%) | (0.01%)
Change in valuation allowance | (41.82%) | (41.29%) | (45.17%)
Deferred Tax Assets |  |  |
Net operating loss carryforwards | $ 35,042 | $ 22,577 |
Tax credit carryforwards | 3,762 | 2,748 |
Capitalized research and development | 4,087 | 4,780 |
Capitalized legal expenses | 1,464 | 1,223 |
Other differences | 815 | 195 |
Total deferred tax assets | 45,170 | 31,523 |
Valuation allowance | (45,170) | (31,523) |
Valuation allowance |  |  |
Recorded increase in valuation allowance | 13,600 | $ 9,000 | $ 8,200
Excess tax benefit from exercise of stock options included in federal and state net operating loss | 400 |  |
Federal |  |  |
Valuation allowance |  |  |
Net operating loss carryforwards | 89,800 |  |
Tax credit carryforwards | 3,000 |  |
Federal | Additional Paid-in Capital |  |  |
Valuation allowance |  |  |
Net operating loss carryforwards | 400 |  |
State |  |  |
Valuation allowance |  |  |
Net operating loss carryforwards | 88,400 |  |
Tax credit carryforwards | 1,200 |  |
State | Additional Paid-in Capital |  |  |
Valuation allowance |  |  |
Net operating loss carryforwards | $ 400 |  |

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