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Condensed Consolidated Statements of Operations - USD ($)
$ in Thousands
 3 Months Ended 6 Months Ended
Jun. 30, 2017
Jun. 30, 2016
Jun. 30, 2017
Jun. 30, 2016
Operating expenses:        
Research and development $ 4,519 $ 6,818 $ 9,917 $ 13,254
General and administrative 2,400 2,578 4,763 5,348
Total operating expenses 6,919 9,396 14,680 18,602
Loss from operations (6,919) (9,396) (14,680) (18,602)
Other (expense) income:        
Interest expense (127) (220) (276) (463)
Interest and investment income 44 80 83 133
Other income, net 28 91 23 69
Total other expense, net (55) (49) (170) (261)
Net loss $ (6,974) $ (9,445) $ (14,850) $ (18,863)
Net loss per share - basic and diluted (in dollars per share) $ (0.32) $ (0.61) $ (0.73) $ (1.23)
Weighted-average common shares outstanding used in net loss per share - basic and diluted (in shares) 21,796,194 15,373,964 20,452,200 15,354,740