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Stock Incentive Plans (Tables) | 6 Months Ended
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Jun. 30, 2021
Stock Incentive Plans |
Summary of stock option activity | ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | ​ | ​ | ​ | ​ | Weighted | ​ | ​ | ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | Average | ​ | ​ | ​
​ | ​ | ​ | ​ | Weighted- | ​ | Remaining | ​ | Aggregate
​ | ​ | ​ | ​ | Average Exercise | ​ | Contractual | ​ | Intrinsic Value
​ |  | Shares |  | Price |  | Term (years) |  | (in thousands)
Outstanding at December 31, 2020 |  | 1,367,667 | ​ | $ | 11.38 |  | 8.13 | ​ | $ | —
Granted |  | 6,447,000 | ​ | $ | 2.74 | ​ | ​ | ​ | ​ | ​
Assumed in Quellis Acquisition | ​ | 332,494 | ​ | $ | 0.29 | ​ | ​ | ​ | ​ | ​
Cancelled or forfeited |  | (208,152) | ​ | $ | 8.71 | ​ | ​ | ​ | ​ | ​
Expired | ​ | (8,675) | ​ | $ | 16.70 | ​ | ​ | ​ | ​ | ​
Outstanding at June 30, 2021 | ​ | 7,930,334 | ​ | $ | 3.96 | ​ | 9.44 | ​ | $ | 662
Vested and exercisable at June 30, 2021 |  | 770,141 | ​ | $ | 14.07 | ​ | 7.31 | ​ | $ | 207

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