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Stock Incentive Plans (Tables) | 9 Months Ended
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Sep. 30, 2021
Stock Incentive Plans |
Summary of stock option activity | ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | ​ | ​ | ​ | ​ | Weighted | ​ | ​ | ​
​ | ​ | ​ | ​ | ​ | ​ | Average | ​ | ​ | ​
​ | ​ | ​ | ​ | Weighted- | ​ | Remaining | ​ | Aggregate
​ | ​ | ​ | ​ | Average | ​ | Contractual | ​ | Intrinsic Value
​ |  | Shares |  | Exercise Price |  | Term (years) |  | (in thousands)
Outstanding at December 31, 2020 |  | 227,846 | ​ | $ | 68.25 |  | 8.13 | ​ | $ | —
Granted |  | 1,113,982 | ​ | $ | 16.21 | ​ | ​ | ​ | ​ | ​
Assumed in Quellis Acquisition | ​ | 55,414 | ​ | $ | 1.73 | ​ | ​ | ​ | ​ | ​
Exercised | ​ | (2,992) | ​ | $ | 2.10 | ​ | ​ | ​ | ​ | ​
Cancelled or forfeited |  | (58,344) | ​ | $ | 62.08 | ​ | ​ | ​ | ​ | ​
Expired | ​ | (1,507) | ​ | $ | 100.85 | ​ | ​ | ​ | ​ | ​
Outstanding at September 30, 2021 | ​ | 1,334,399 | ​ | $ | 22.43 | ​ | 9.25 | ​ | $ | 374
Vested and exercisable at September 30, 2021 |  | 127,064 | ​ | $ | 74.71 | ​ | 7.38 | ​ | $ | 178

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