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Stock Incentive Plans | 9 Months Ended
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Sep. 30, 2021
Stock Incentive Plans |
Stock Incentive Plans | 9. Stock Incentive Plans A summary of the Company’s stock option activity and related information follows: ​
​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​ | ​
​ | ​ | ​ | ​ | ​ | ​ | Weighted | ​ | ​ | ​
​ | ​ | ​ | ​ | ​ | ​ | Average | ​ | ​ | ​
​ | ​ | ​ | ​ | Weighted- | ​ | Remaining | ​ | Aggregate
​ | ​ | ​ | ​ | Average | ​ | Contractual | ​ | Intrinsic Value
​ |  | Shares |  | Exercise Price |  | Term (years) |  | (in thousands)
Outstanding at December 31, 2020 |  | 227,846 | ​ | $ | 68.25 |  | 8.13 | ​ | $ | —
Granted |  | 1,113,982 | ​ | $ | 16.21 | ​ | ​ | ​ | ​ | ​
Assumed in Quellis Acquisition | ​ | 55,414 | ​ | $ | 1.73 | ​ | ​ | ​ | ​ | ​
Exercised | ​ | (2,992) | ​ | $ | 2.10 | ​ | ​ | ​ | ​ | ​
Cancelled or forfeited |  | (58,344) | ​ | $ | 62.08 | ​ | ​ | ​ | ​ | ​
Expired | ​ | (1,507) | ​ | $ | 100.85 | ​ | ​ | ​ | ​ | ​
Outstanding at September 30, 2021 | ​ | 1,334,399 | ​ | $ | 22.43 | ​ | 9.25 | ​ | $ | 374
Vested and exercisable at September 30, 2021 |  | 127,064 | ​ | $ | 74.71 | ​ | 7.38 | ​ | $ | 178
​ The total intrinsic value of options exercised in the three and nine months ended September 30, 2021 was $38 thousand. There were no options exercised in the three or nine months ended September 30, 2020. The total grant date fair value of options vested for the three months ended September 30, 2021 and 2020 was $0.3 million. The total grant date fair value of options vested for the nine months ended September 30, 2021 and 2020 was $1.3 million and $1.0 million, respectively. The weighted-average grant date fair value of options granted to employees and non-employees for the three months ended September 30, 2021 and 2020 was $6.20 and $29.10, respectively. The weighted-average grant date fair value of options granted to employees and non-employees for the nine months ended September 30, 2021 and 2020 was $10.15 and $22.32, respectively. At September 30, 2021, the total unrecognized compensation expense related to unvested stock option awards was $11.8 million. The Company expects to recognize that cost over a weighted-average period of approximately 3.1 years.

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