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Condensed Consolidated Statements of Stockholders' Equity - USD ($) $ in Thousands | Preferred Stock Series X redeemable convertible preferred stock | Common stock | Additional Paid-in Capital | Deficit Accumulated during the Development Stage | Accumulated other comprehensive loss | Total
--- | --- | --- | --- | --- | --- | ---
Balance, beginning of period at Dec. 31, 2019 | $ 0 | $ 2 | $ 259,315 | $ (223,597) | $ 0 |
Balance, beginning of period (in shares) at Dec. 31, 2019 | 0 | 2,072,266 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock in an underwritten public offering, net of issuance costs |  |  | 24,558 |  |  |
Issuance of common stock for at-the-market offerings |  |  | 16,268 |  |  |
Issuance of common stock for at-the-market offerings (in shares) |  | 392,288 |  |  |  |
Issuance of common stock in underwritten public offerings |  | $ 1 |  |  |  |
Issuance of common stock and warrants in public offerings (in shares) |  | 881,666 |  |  |  |
Stock-based compensation expense |  |  | 1,085 |  |  |
Net loss |  |  |  | (28,322) |  | $ (28,322)
Balance, end of period at Sep. 30, 2020 |  | $ 3 | 301,226 | (251,919) | 0 | 49,310
Balance, end of period (in shares) at Sep. 30, 2020 |  | 3,346,220 |  |  |  |
Balance, beginning of period at Jun. 30, 2020 | $ 0 | $ 3 | 291,912 | (241,057) | 1 |
Balance, beginning of period (in shares) at Jun. 30, 2020 | 0 | 3,137,264 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock for at-the-market offerings |  |  | 8,939 |  |  |
Issuance of common stock for at-the-market offerings (in shares) |  | 208,956 |  |  |  |
Stock-based compensation expense |  |  | 375 |  |  |
Net loss |  |  |  | (10,862) |  | (10,862)
Realized loss on short-term investments |  |  |  |  | (1) | (1)
Balance, end of period at Sep. 30, 2020 |  | $ 3 | 301,226 | (251,919) | $ 0 | 49,310
Balance, end of period (in shares) at Sep. 30, 2020 |  | 3,346,220 |  |  |  |
Balance, beginning of period at Dec. 31, 2020 | $ 0 | $ 4 | 301,562 | (260,897) |  | $ 40,669
Balance, beginning of period (in shares) at Dec. 31, 2020 | 0 | 3,347,386 |  |  |  | 3,347,389
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Reclassification of preferred stock to permanent equity | $ 240,881 |  |  |  |  |
Reclassification of preferred stock to permanent equity (in shares) | 86,077 |  |  |  |  |
Conversion of preferred stock into common stock | $ (168,920) |  |  |  |  |
Conversion of preferred stock into common stock (in shares) | (54,622) |  |  |  |  |
Issuance of preferred stock in a private offering of public equity, net of issuance costs |  |  | 19,565 |  |  |
Issuance of common stock upon acquisition of Quellis |  |  | 8,098 |  |  |
Issuance of common stock upon acquisition of Quellis (in shares) |  | 555,444 |  |  |  |
Issuance of common stock upon the conversion of preferred stock |  | $ 9 | 168,912 |  |  |
Issuance of common stock upon the conversion of preferred stock (in shares) |  | 9,103,664 |  |  |  |
Dividend on convertible preferred stock related to beneficial conversion feature and issuance costs | $ 24,437 |  | (24,437) |  |  |
Expense related to warrants inherited in acquisition of Quellis |  |  | 772 |  |  |
Reclassification of warrant liability to additional paid-in capital |  |  | 3,468 |  |  |
Issuance of common stock upon exercise of options |  |  | 6 |  |  |
Fractional shares eliminated pursuant to reverse stock split |  | (9) |  |  |  |
Stock-based compensation expense |  |  | 2,390 |  |  |
Net loss |  |  |  | (185,421) |  | $ (185,421)
Balance, end of period at Sep. 30, 2021 | $ 96,398 | $ 13 | 480,336 | (446,318) |  | $ 130,429
Balance, end of period (in shares) at Sep. 30, 2021 | 31,455 | 13,009,477 |  |  |  | 13,009,477
Balance, beginning of period at Jun. 30, 2021 | $ 99,770 | $ 13 | 475,563 | (438,447) |  |
Balance, beginning of period (in shares) at Jun. 30, 2021 | 32,545 | 12,824,796 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Conversion of preferred stock into common stock | $ (3,372) |  |  |  |  |
Conversion of preferred stock into common stock (in shares) | (1,090) |  |  |  |  |
Issuance of common stock upon the conversion of preferred stock |  |  | 3,372 |  |  |
Issuance of common stock upon the conversion of preferred stock (in shares) |  | 181,698 |  |  |  |
Expense related to warrants inherited in acquisition of Quellis |  |  | 385 |  |  |
Issuance of common stock upon exercise of options |  |  | 6 |  |  |
Fractional shares eliminated pursuant to reverse stock split |  | (9) |  |  |  |
Stock-based compensation expense |  |  | 1,010 |  |  |
Net loss |  |  |  | (7,871) |  | $ (7,871)
Balance, end of period at Sep. 30, 2021 | $ 96,398 | $ 13 | $ 480,336 | $ (446,318) |  | $ 130,429
Balance, end of period (in shares) at Sep. 30, 2021 | 31,455 | 13,009,477 |  |  |  | 13,009,477

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