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Organization and Operations - Stock Purchase Agreement and Series X Preferred Stock (Details) $ in Thousands |  | 1 Months Ended | 3 Months Ended | 6 Months Ended |  |
--- | --- | --- | --- | --- | --- | ---
Jan. 28, 2021 USD ($) shares | Feb. 28, 2021 shares | Jun. 30, 2021 shares | Mar. 31, 2021 USD ($) shares | Jun. 30, 2022 shares | Jun. 30, 2021 USD ($) | Dec. 31, 2021 shares | Jun. 02, 2021
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Net proceeds from issuance of private placement | $ |  |  |  |  |  | $ 104,261 |  |
Beneficial conversion feature | $ |  |  |  | $ 19,565 |  |  |  |
Common Stock Issuable Upon Conversion at Transaction Date |  | 14,813,954 |  |  |  |  |  |
Minimum |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Common stock, percentage of beneficial ownership initially, after conversion |  |  |  |  | 4.99% |  |  |
Common stock, percentage of beneficial ownership thereafter, after conversion |  |  |  |  | 4.99% |  |  |
Maximum |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Common stock, percentage of beneficial ownership initially, after conversion |  |  |  |  | 9.99% |  |  |
Common stock, percentage of beneficial ownership thereafter, after conversion |  |  |  |  | 19.99% |  |  |
Shares issued in merger |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Common Stock Issuable Upon Conversion at Transaction Date |  | 8,417,502 |  |  |  |  |  |
Shares issued in February 2021 Financing |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Common Stock Issuable Upon Conversion at Transaction Date |  | 5,928,952 |  |  |  |  |  |
Warrants assumed in merger |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Common Stock Issuable Upon Conversion at Transaction Date |  | 467,500 |  |  |  |  |  |
Series X redeemable convertible preferred stock |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Issuance of preferred stock in a private offering of public equity, net of issuance costs (in shares) |  |  |  | 35,573 |  |  |  |
Number of shares of common stock into which each share of Series X Preferred Stock may be converted |  |  |  |  | 166.67 |  |  |
Series X Preferred Stock at Transaction Date |  | 88,882 |  |  |  |  |  |
Common Stock Issuable Upon Conversion at Transaction Date |  |  |  |  | 5,242,501 |  |  |
Number of preferred shares converted during period |  |  |  |  | 54,622 |  |  |
Preferred stock, outstanding (in shares) |  |  |  |  | 31,455 |  | 31,455 |
Series X redeemable convertible preferred stock | Shares issued in merger |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Series X Preferred Stock at Transaction Date |  | 50,504 |  |  |  |  |  |
Series X redeemable convertible preferred stock | Shares issued in February 2021 Financing |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Series X Preferred Stock at Transaction Date |  | 35,573 |  |  |  |  |  |
Series X redeemable convertible preferred stock | Warrants assumed in merger |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Series X Preferred Stock at Transaction Date |  | 2,805 |  |  |  |  |  |
Common stock |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Number of shares of common stock into which each share of Series X Preferred Stock may be converted |  |  |  |  | 166.67 |  |  | 166.67
Common Stock Issuable Upon Conversion at Transaction Date |  |  |  |  | 5,242,501 |  |  |
Number of common shares issued from conversion of Series X preferred shares |  |  | 8,921,966 |  | 9,103,664 |  |  |
Quellis Biosciences, Inc | Stock Purchase Agreement |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Beneficial conversion feature | $ | $ 19,600 |  |  |  |  |  |  |
Quellis Biosciences, Inc | Series X redeemable convertible preferred stock |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Number of shares of common stock into which each share of Series X Preferred Stock may be converted | 166.67 |  |  |  |  |  |  |
Quellis Biosciences, Inc | Series X redeemable convertible preferred stock | Stock Purchase Agreement |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Issuance of preferred stock in a private offering of public equity, net of issuance costs (in shares) | 35,573 |  |  |  |  |  |  |
Gross proceeds from issuance of preferred stock in a private offering | $ | $ 110,000 |  |  |  |  |  |  |
Net proceeds from issuance of private placement | $ | 104,300 |  |  |  |  |  |  |
Issuance costs | $ | $ 5,700 |  |  |  |  |  |  |
Quellis Biosciences, Inc | Common stock | Series X redeemable convertible preferred stock |  |  |  |  |  |  |  |
Stock Purchase Agreement and Series X Preferred Stock |  |  |  |  |  |  |  |
Number of shares of common stock into which each share of Series X Preferred Stock may be converted | 166.67 |  |  |  |  |  |  |

X
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