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Condensed Consolidated Statements of Redeemable Convertible Preferred Stock and Stockholders' Equity - USD ($) $ in Thousands | Preferred stock Series X redeemable convertible preferred stock | Common stock | Additional paid-in capital | Accumulated deficit | Accumulated other comprehensive loss | Total
--- | --- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2020 | $ 0 | $ 4 | $ 301,562 | $ (260,897) | $ 0 | $ 40,669
Beginning balance (in shares) at Dec. 31, 2020 | 0 | 3,347,386 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of preferred stock in a private offering of public equity, net of issuance costs |  |  | 19,565 |  |  | 19,565
Issuance of preferred stock and common stock upon acquisition of Quellis |  |  | 8,098 |  |  | 8,098
Issuance of preferred stock and common stock upon acquisition of Quellis(in shares) |  | 555,444 |  |  |  |
Expense related to warrants inherited in acquisition of Quellis |  |  | 241 |  |  | 241
Stock-based compensation expense |  |  | 366 |  |  | 366
Net loss |  |  |  | (170,084) |  | (170,084)
Ending balance at Mar. 31, 2021 |  | $ 4 | 329,832 | (430,981) |  | (101,145)
Ending balance (in shares) at Mar. 31, 2021 |  | 3,902,830 |  |  |  |
Beginning balance at Dec. 31, 2020 | $ 0 | $ 4 | 301,562 | (260,897) | 0 | 40,669
Beginning balance (in shares) at Dec. 31, 2020 | 0 | 3,347,386 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Net loss |  |  |  |  |  | (177,550)
Ending balance at Jun. 30, 2021 | $ 99,770 | $ 13 | 475,563 | (438,447) | 0 | 136,899
Ending balance (in shares) at Jun. 30, 2021 | 32,545 | 12,824,796 |  |  |  |
Beginning balance at Mar. 31, 2021 |  | $ 4 | 329,832 | (430,981) |  | (101,145)
Beginning balance (in shares) at Mar. 31, 2021 |  | 3,902,830 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Reclassification of preferred stock to permanent equity | $ 240,881 |  |  |  |  | 240,881
Reclassification of preferred stock to permanent equity (in shares) | 86,077 |  |  |  |  |
Issuance of common stock upon the conversion of preferred stock | $ (165,548) | $ 9 | 165,540 |  |  | 1
Issuance of common stock upon the conversion of preferred stock (in shares) | (53,532) | 8,921,966 |  |  |  |
Accretion of preferred stock discount | $ 24,437 |  | (24,437) |  |  |
Expense related to warrants inherited in acquisition of Quellis |  |  | 146 |  |  | 146
Reclassification of equity classified warrants |  |  | 3,468 |  |  | 3,468
Stock-based compensation expense |  |  | 1,014 |  |  | 1,014
Net loss |  |  |  | (7,466) |  | (7,466)
Ending balance at Jun. 30, 2021 | $ 99,770 | $ 13 | 475,563 | (438,447) | 0 | 136,899
Ending balance (in shares) at Jun. 30, 2021 | 32,545 | 12,824,796 |  |  |  |
Beginning balance at Dec. 31, 2021 | $ 96,398 | $ 13 | 481,709 | (455,809) | 0 | 122,311
Beginning balance (in shares) at Dec. 31, 2021 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Expense related to warrants inherited in acquisition of Quellis |  |  | 1,542 |  |  | 1,542
Stock-based compensation expense |  |  | 1,209 |  |  | 1,209
Net loss |  |  |  | (15,324) |  | (15,324)
Unrealized loss on short-term investments |  |  |  |  | (53) | (53)
Ending balance at Mar. 31, 2022 | $ 96,398 | $ 13 | 484,460 | (471,133) | (53) | 109,685
Ending balance (in shares) at Mar. 31, 2022 | 31,455 | 13,016,955 |  |  |  |
Beginning balance at Dec. 31, 2021 | $ 96,398 | $ 13 | 481,709 | (455,809) | 0 | 122,311
Beginning balance (in shares) at Dec. 31, 2021 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock upon the conversion of preferred stock (in shares) |  | 9,103,664 |  |  |  |
Net loss |  |  |  |  |  | (26,574)
Realized loss on short-term investments |  |  |  |  |  | (186)
Ending balance at Jun. 30, 2022 | $ 96,398 | $ 13 | 485,577 | (482,383) | (186) | 99,419
Ending balance (in shares) at Jun. 30, 2022 | 31,455 | 13,016,955 |  |  |  |
Beginning balance at Mar. 31, 2022 | $ 96,398 | $ 13 | 484,460 | (471,133) | (53) | 109,685
Beginning balance (in shares) at Mar. 31, 2022 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Stock-based compensation expense |  |  | 1,117 |  |  | 1,117
Net loss |  |  |  | (11,250) |  | (11,250)
Unrealized loss on short-term investments |  |  |  |  | (133) | (133)
Realized loss on short-term investments |  |  |  |  |  | (133)
Ending balance at Jun. 30, 2022 | $ 96,398 | $ 13 | $ 485,577 | $ (482,383) | $ (186) | $ 99,419
Ending balance (in shares) at Jun. 30, 2022 | 31,455 | 13,016,955 |  |  |  |

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