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Condensed Consolidated Statements of Redeemable Convertible Preferred Stock and Stockholders' Equity - USD ($) $ in Thousands | Preferred stock Series X redeemable convertible preferred stock | Common stock | Additional paid-in capital | Accumulated deficit | Accumulated other comprehensive loss | Total
--- | --- | --- | --- | --- | --- | ---
Beginning balance at Dec. 31, 2021 | $ 96,398 | $ 13 | $ 481,709 | $ (455,809) | $ 0 | $ 122,311
Beginning balance (in shares) at Dec. 31, 2021 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Expense related to warrants inherited in acquisition of Quellis |  |  | 1,542 |  |  | 1,542
Stock-based compensation expense |  |  | 1,209 |  |  | 1,209
Unrealized gain (loss) on short-term investments |  |  |  |  | (53) | (53)
Net loss |  |  |  | (15,324) |  | (15,324)
Ending balance at Mar. 31, 2022 | $ 96,398 | $ 13 | 484,460 | (471,133) | (53) | 109,685
Ending balance (in shares) at Mar. 31, 2022 | 31,455 | 13,016,955 |  |  |  |
Beginning balance at Dec. 31, 2021 | $ 96,398 | $ 13 | 481,709 | (455,809) | 0 | 122,311
Beginning balance (in shares) at Dec. 31, 2021 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Net loss |  |  |  |  |  | (38,571)
Ending balance at Sep. 30, 2022 | $ 96,398 | $ 16 | 511,053 | (494,380) | (215) | 112,872
Ending balance (in shares) at Sep. 30, 2022 | 31,455 | 15,748,323 |  |  |  |
Beginning balance at Mar. 31, 2022 | $ 96,398 | $ 13 | 484,460 | (471,133) | (53) | 109,685
Beginning balance (in shares) at Mar. 31, 2022 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Stock-based compensation expense |  |  | 1,117 |  |  | 1,117
Unrealized gain (loss) on short-term investments |  |  |  |  | (133) | (133)
Net loss |  |  |  | (11,250) |  | (11,250)
Ending balance at Jun. 30, 2022 | $ 96,398 | $ 13 | 485,577 | (482,383) | (186) | 99,419
Ending balance (in shares) at Jun. 30, 2022 | 31,455 | 13,016,955 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock upon exercise of options |  |  | 34 |  |  | 34
Issuance of common stock upon exercise of options (in shares) |  | 16,202 |  |  |  |
Issuance of common stock for at-the-market offerings, net of issuance costs |  | $ 3 | 24,287 |  |  | 24,290
Issuance of common stock for at-the-market offerings, net of issuance costs (in shares) |  | 2,715,166 |  |  |  |
Stock-based compensation expense |  |  | 1,155 |  |  | 1,155
Unrealized gain (loss) on short-term investments |  |  |  |  | (29) | (29)
Net loss |  |  |  | (11,997) |  | (11,997)
Ending balance at Sep. 30, 2022 | $ 96,398 | $ 16 | 511,053 | (494,380) | (215) | 112,872
Ending balance (in shares) at Sep. 30, 2022 | 31,455 | 15,748,323 |  |  |  |
Beginning balance at Dec. 31, 2022 | $ 96,398 | $ 28 | 632,512 | (507,643) | (79) | 221,216
Beginning balance (in shares) at Dec. 31, 2022 | 31,455 | 27,501,340 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock upon the conversion of preferred stock | $ (1,074) |  | 1,074 |  |  |
Issuance of common stock upon the conversion of preferred stock (shares) | (348) | 57,910 |  |  |  |
Issuance of common stock upon exercise of options and warrants |  |  | 37 |  |  | 37
Issuance of common stock upon exercise of options and warrants (in shares) |  | 427,468 |  |  |  |
Stock-based compensation expense |  |  | 1,220 |  |  | 1,220
Unrealized gain (loss) on short-term investments |  |  |  |  | 75 | 75
Net loss |  |  |  | (11,188) |  | (11,188)
Ending balance at Mar. 31, 2023 | $ 95,324 | $ 28 | 634,843 | (518,831) | (4) | 211,360
Ending balance (in shares) at Mar. 31, 2023 | 31,107 | 27,986,718 |  |  |  |
Beginning balance at Dec. 31, 2022 | $ 96,398 | $ 28 | 632,512 | (507,643) | (79) | 221,216
Beginning balance (in shares) at Dec. 31, 2022 | 31,455 | 27,501,340 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Net loss |  |  |  |  |  | (41,481)
Ending balance at Sep. 30, 2023 | $ 95,324 | $ 28 | 638,251 | (549,124) |  | 184,479
Ending balance (in shares) at Sep. 30, 2023 | 31,107 | 28,042,296 |  |  |  |
Beginning balance at Mar. 31, 2023 | $ 95,324 | $ 28 | 634,843 | (518,831) | (4) | 211,360
Beginning balance (in shares) at Mar. 31, 2023 | 31,107 | 27,986,718 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock upon exercise of options |  |  | 273 |  |  | 273
Issuance of common stock upon exercise of options (in shares) |  | 39,126 |  |  |  |
Stock-based compensation expense |  |  | 1,331 |  |  | 1,331
Unrealized gain (loss) on short-term investments |  |  |  |  | $ 4 | 4
Net loss |  |  |  | (12,566) |  | (12,566)
Ending balance at Jun. 30, 2023 | $ 95,324 | $ 28 | 636,447 | (531,397) |  | 200,402
Ending balance (in shares) at Jun. 30, 2023 | 31,107 | 28,025,844 |  |  |  |
Increase (Decrease) in Shareholders' Equity |  |  |  |  |  |
Issuance of common stock upon exercise of options |  |  | 110 |  |  | 110
Issuance of common stock upon exercise of options (in shares) |  | 16,452 |  |  |  |
Stock-based compensation expense |  |  | 1,694 |  |  | 1,694
Net loss |  |  |  | (17,727) |  | (17,727)
Ending balance at Sep. 30, 2023 | $ 95,324 | $ 28 | $ 638,251 | $ (549,124) |  | $ 184,479
Ending balance (in shares) at Sep. 30, 2023 | 31,107 | 28,042,296 |  |  |  |

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