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Condensed Consolidated Statements of Redeemable Convertible Preferred Stock and Stockholders' Equity - USD ($)
$ in Thousands
Preferred stock
Series X redeemable convertible preferred stock
Common stock
Additional paid-in capital
Accumulated deficit
Accumulated other comprehensive loss
Total
Beginning balance at Dec. 31, 2021 $ 96,398 $ 13 $ 481,709 $ (455,809) $ 0 $ 122,311
Beginning balance (in shares) at Dec. 31, 2021 31,455 13,016,955        
Increase (Decrease) in Shareholders' Equity            
Expense related to warrants inherited in acquisition of Quellis     1,542     1,542
Stock-based compensation expense     1,209     1,209
Unrealized gain (loss) on short-term investments         (53) (53)
Net loss       (15,324)   (15,324)
Ending balance at Mar. 31, 2022 $ 96,398 $ 13 484,460 (471,133) (53) 109,685
Ending balance (in shares) at Mar. 31, 2022 31,455 13,016,955        
Beginning balance at Dec. 31, 2021 $ 96,398 $ 13 481,709 (455,809) 0 122,311
Beginning balance (in shares) at Dec. 31, 2021 31,455 13,016,955        
Increase (Decrease) in Shareholders' Equity            
Net loss           (38,571)
Ending balance at Sep. 30, 2022 $ 96,398 $ 16 511,053 (494,380) (215) 112,872
Ending balance (in shares) at Sep. 30, 2022 31,455 15,748,323        
Beginning balance at Mar. 31, 2022 $ 96,398 $ 13 484,460 (471,133) (53) 109,685
Beginning balance (in shares) at Mar. 31, 2022 31,455 13,016,955        
Increase (Decrease) in Shareholders' Equity            
Stock-based compensation expense     1,117     1,117
Unrealized gain (loss) on short-term investments         (133) (133)
Net loss       (11,250)   (11,250)
Ending balance at Jun. 30, 2022 $ 96,398 $ 13 485,577 (482,383) (186) 99,419
Ending balance (in shares) at Jun. 30, 2022 31,455 13,016,955        
Increase (Decrease) in Shareholders' Equity            
Issuance of common stock upon exercise of options     34     34
Issuance of common stock upon exercise of options (in shares)   16,202        
Issuance of common stock for at-the-market offerings, net of issuance costs   $ 3 24,287     24,290
Issuance of common stock for at-the-market offerings, net of issuance costs (in shares)   2,715,166        
Stock-based compensation expense     1,155     1,155
Unrealized gain (loss) on short-term investments         (29) (29)
Net loss       (11,997)   (11,997)
Ending balance at Sep. 30, 2022 $ 96,398 $ 16 511,053 (494,380) (215) 112,872
Ending balance (in shares) at Sep. 30, 2022 31,455 15,748,323        
Beginning balance at Dec. 31, 2022 $ 96,398 $ 28 632,512 (507,643) (79) 221,216
Beginning balance (in shares) at Dec. 31, 2022 31,455 27,501,340        
Increase (Decrease) in Shareholders' Equity            
Issuance of common stock upon the conversion of preferred stock $ (1,074)   1,074      
Issuance of common stock upon the conversion of preferred stock (shares) (348) 57,910        
Issuance of common stock upon exercise of options and warrants     37     37
Issuance of common stock upon exercise of options and warrants (in shares)   427,468        
Stock-based compensation expense     1,220     1,220
Unrealized gain (loss) on short-term investments         75 75
Net loss       (11,188)   (11,188)
Ending balance at Mar. 31, 2023 $ 95,324 $ 28 634,843 (518,831) (4) 211,360
Ending balance (in shares) at Mar. 31, 2023 31,107 27,986,718        
Beginning balance at Dec. 31, 2022 $ 96,398 $ 28 632,512 (507,643) (79) 221,216
Beginning balance (in shares) at Dec. 31, 2022 31,455 27,501,340        
Increase (Decrease) in Shareholders' Equity            
Net loss           (41,481)
Ending balance at Sep. 30, 2023 $ 95,324 $ 28 638,251 (549,124)   184,479
Ending balance (in shares) at Sep. 30, 2023 31,107 28,042,296        
Beginning balance at Mar. 31, 2023 $ 95,324 $ 28 634,843 (518,831) (4) 211,360
Beginning balance (in shares) at Mar. 31, 2023 31,107 27,986,718        
Increase (Decrease) in Shareholders' Equity            
Issuance of common stock upon exercise of options     273     273
Issuance of common stock upon exercise of options (in shares)   39,126        
Stock-based compensation expense     1,331     1,331
Unrealized gain (loss) on short-term investments         $ 4 4
Net loss       (12,566)   (12,566)
Ending balance at Jun. 30, 2023 $ 95,324 $ 28 636,447 (531,397)   200,402
Ending balance (in shares) at Jun. 30, 2023 31,107 28,025,844        
Increase (Decrease) in Shareholders' Equity            
Issuance of common stock upon exercise of options     110     110
Issuance of common stock upon exercise of options (in shares)   16,452        
Stock-based compensation expense     1,694     1,694
Net loss       (17,727)   (17,727)
Ending balance at Sep. 30, 2023 $ 95,324 $ 28 $ 638,251 $ (549,124)   $ 184,479
Ending balance (in shares) at Sep. 30, 2023 31,107 28,042,296