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Collaborative, Licensing and Other Arrangements - NIAID - Additional Information (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended
--- | --- | ---
Oct. 31, 2011 | Sep. 30, 2008 | Dec. 31, 2015 | Dec. 31, 2014 | Dec. 31, 2013 | Dec. 31, 2010 | Dec. 31, 2009
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Deferred revenue |  |  |  | $ 1,939 |  |  |
Recognition of revenue |  |  | $ 6,383 | 13,183 | $ 24,423 |  |
National Institute of Allergy and Infectious Diseases "NIAID" [Member] | Arrangement with Governmental Agency 2 [Member] |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Total contract amount awarded |  | $ 64,800 |  |  |  |  |
Increase (decrease) in revenue |  |  |  |  |  | $ 900 | $ 1,100
Recognition of revenue |  |  | 200 | 1,200 | 4,400 |  |
National Institute of Allergy and Infectious Diseases "NIAID" [Member] | Arrangement with Governmental Agency 3 [Member] |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Contractual term |  | 3 years |  |  |  |  |
Increase (decrease) in deferred revenue |  |  |  | (1,800) |  |  |
Deferred revenue |  |  |  | 1,900 |  |  |
Remaining deferred revenue |  |  |  | 100 |  |  |
National Institute of Allergy and Infectious Diseases "NIAID" [Member] | Arrangement with Governmental Agency 1 [Member] |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Total contract amount awarded | $ 28,000 |  |  |  |  |  |
Contractual term | 5 years |  |  |  |  |  |
Recognition of revenue |  |  | $ 4,900 | $ 8,400 | $ 4,700 |  |

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- Definition Revenue earned during the period arising from products sold or services provided under the terms of a contract, not elsewhere specified in the taxonomy. May include government contracts, construction contracts, and any other contract related to a particular project or product. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 225 -SubTopic 10 -Section S99 -Paragraph 2 -Subparagraph (SX 210.5-03.1) -URI http://asc.fasb.org/extlink&oid=26872669&loc=d3e20235-122688 + Details
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