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Collaborative, Licensing and Other Arrangements - Servier - Additional Information (Details) $ in Thousands, € in Millions | 1 Months Ended | 12 Months Ended
--- | --- | ---
Jan. 31, 2011 USD ($) | Jan. 31, 2011 EUR (€) | Dec. 31, 2015 USD ($) | Dec. 31, 2015 EUR (€) | Dec. 31, 2014 USD ($) | Dec. 31, 2013 USD ($) | Dec. 31, 2011
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Deferred revenue – current |  |  | $ 3,198 |  | $ 1,089 |  |
Collaborative Arrangement [Member] | Servier [Member] |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |
Upfront payment received | $ 15,000 |  |  |  |  |  |
Period for recognizing deferred revenue |  |  |  |  |  |  | 8 months
Proceeds from loan | $ 19,500 | € 15.0 |  |  |  |  |
Future initial research and development expenses to be funded by counterparty |  |  | 50,000 |  |  |  |
Contract and other revenue |  |  | 1,200 |  | $ 3,500 | $ 13,600 |
Eligible milestone payments receivable | € |  |  |  | € 356.5 |  |  |
Eligible milestone payments receivable under specific rights not met | € |  |  |  | 633.8 |  |  |
Eligible milestone payments receivable, after amendment | € |  |  |  | 341.5 |  |  |
Eligible milestone payments receivable under specific rights not met, after amendment | € |  |  |  | € 618.8 |  |  |
Deferred revenue – current |  |  | $ 600 |  |  |  |

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