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Collaborative, Licensing and Other Arrangements - Pfizer - Additional Information (Details) - Pfizer Inc. [Member] - USD ($) $ in Millions |  | 1 Months Ended | 12 Months Ended
--- | --- | --- | ---
Dec. 03, 2015 | Aug. 31, 2005 | Dec. 31, 2015 | Dec. 31, 2007
Collaborative Arrangement [Member] |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |
Milestone received under the collaboration agreement | $ 3.8 |  | $ 4.2 |
Royalty right expiration period |  | 10 years |  |
License Arrangement [Member] |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |
Proceeds from license fees received |  |  |  | $ 30.0

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