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Collaborative and Other Agreements - Servier - Additional Information (Details) - Collaborative Arrangement [Member] - Servier [Member] - USD ($) | 3 Months Ended | 6 Months Ended
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Jun. 30, 2016 | Jun. 30, 2015 | Jun. 30, 2016 | Mar. 25, 2016 | Jun. 30, 2015
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |
Future initial research and development expenses to be funded by counterparty |  |  | $ 50,000,000 |  |
Contract and other revenue | $ 0 | $ 300,000 | $ 300,000 |  | $ 900,000
Deferred revenue recognized |  |  |  | $ 600,000 |

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- Definition Amount of revenue recognized that was previously reported as deferred or unearned revenue. + References Reference 1: http://www.xbrl.org/2003/role/presentationRef -Publisher FASB -Name Accounting Standards Codification -Topic 605 -SubTopic 10 -Section S99 -Paragraph 1 -Subparagraph (SAB TOPIC 13.A.4(a).Q1 Response)) -URI http://asc.fasb.org/extlink&oid=65893310&loc=d3e214044-122780 + Details
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