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Collaborative and Other Agreements - NIAID - Additional Information (Details) - USD ($) | 1 Months Ended | 3 Months Ended | 9 Months Ended
--- | --- | --- | ---
Oct. 31, 2011 | Sep. 30, 2016 | Sep. 30, 2015 | Sep. 30, 2016 | Sep. 30, 2015
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |
Recognition of revenue |  | $ 205,000 | $ 1,429,000 | $ 1,844,000 | $ 5,412,000
National Institute of Allergy and Infectious Diseases "NIAID" [Member] |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |
Total contract amount awarded | $ 28,000,000 |  |  |  |
Contractual term | 5 years |  |  |  |
Recognition of revenue |  | $ 0 | $ 1,200,000 | $ 1,100,000 | $ 4,100,000

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