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Collaborative, Licensing and Other Arrangements - Servier - Additional Information (Details) - Collaborative Arrangement [Member] - Servier [Member] - USD ($) | 3 Months Ended | 6 Months Ended
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Jun. 30, 2017 | Jun. 30, 2016 | Jun. 30, 2017 | Jun. 30, 2016 | Mar. 25, 2016
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |
Future initial research and development expenses to be funded by counterparty |  |  | $ 50,000,000 |  |
Contract and other revenue | $ 0 | $ 0 | $ 0 | $ 300,000 |
Deferred revenue recognized |  |  |  |  | $ 600,000

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