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Commitments and Contingencies - Additional Information (Details) |  |  |  | 1 Months Ended | 12 Months Ended
--- | --- | --- | --- | --- | ---
Dec. 31, 2018 USD ($) Facility | Apr. 14, 2018 USD ($) Facility | Nov. 21, 2017 USD ($) Facility | Oct. 31, 2018 USD ($) Facility | Sep. 30, 2017 | Dec. 31, 2018 USD ($) Facility | Dec. 31, 2017 USD ($)
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Estimate of milestone payments | $ 7,500,000 |  |  |  |  | $ 7,500,000 |
Operating leases, expiry date |  |  |  |  |  | 2023-04 |
Operating leases, number of leased facilities | Facility |  | 3 | 3 | 3 |  |  |
Total lease payments | 14,925,000 |  |  |  |  | $ 14,925,000 |
Operating leases, total rental expense |  |  |  |  |  | 2,100,000 | $ 2,400,000
Operating leases, number of portions of leased facilities | Facility |  | 1 | 1 | 1 |  |  |
Operating leases, sublease commencement date |  | May 01, 2018 | Dec. 26, 2017 | Oct. 24, 2018 |  |  |
Operating leases sublease income plus reimbursement of operating expenses receivable |  | $ 1,100,000 | $ 5,100,000 |  |  |  |
Operating leases sublease expiry date |  | 2023-04 | 2023-04 | 2021-05 |  |  |
Sublease tenant improvement allowance |  | $ 65,000 | $ 800,000 |  |  |  |
Operating leases sublease income receivable |  |  |  | $ 1,700,000 |  |  |
Llease Agreement Commenced in Septembr 2017 [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Lease agreement term |  |  |  |  | 63 months |  |
Lease commencement date |  |  |  |  | Nov. 14, 2017 |  |
Total lease payments | 800,000 |  |  |  |  | $ 800,000 |
Lease payments term |  |  |  |  | 2023-02 |  |
Sublease Agreement Commenced on December 26, 2017 [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Sublease loss |  |  | 800,000 |  |  |  |
Sublease income |  |  | 1,500,000 |  |  |  |
Sublease Agreement Commenced on December 26, 2017 [Member] | Standby Letter of Credit [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Letter of credit amount |  |  | $ 1,000,000 |  |  |  |
Funds drawn from letter of credit | $ 0 |  |  |  |  |  |
Sublease Agreement Commenced on May 1,2018 [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Sublease income |  | 300,000 |  |  |  |  |
Sublease broker commissions |  | 89,000 |  |  |  |  |
Sublease Agreement Commenced on May 1,2018 [Member] | Restructuring Charges [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Sublease loss |  | $ 600,000 |  |  |  |  |
Sublease Agreement Commenced on October 24,2018 [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Sublease loss |  |  |  | 0 |  |  |
Sublease broker commissions |  |  |  | $ 137,000 |  |  |
Berkeley, California [Member] |  |  |  |  |  |  |
Commitments And Contingencies [Line Items] |  |  |  |  |  |  |
Operating leases, number of leased facilities | Facility | 2 |  |  |  |  | 2 |

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