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Compensation and Other Benefit Plans (Tables) | 12 Months Ended
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Dec. 31, 2019
Compensation and Other Benefit Plans |
Stock Option Activity | The following table summarizes the Company’s stock option activity for the year ended December 31, 2019:
|  |  |  | Weighted |  | Weighted |  | Aggregate
|  |  |  | Average |  | Average |  | Intrinsic
|  |  |  | Exercise |  | Contractual |  | Value
|  | Number of |  | Price |  | Term |  | (in
|  | shares |  | Per Share |  | (in years) |  | thousands)
Outstanding at beginning of year |  | 1,624,746 |  | $ | 23.09 |  | 7.5 |  | $ | 8,104
Granted |  | 438,814 |  |  | 14.84 |  |  |  |  |
Exercised |  | (55,759) |  |  | 4.90 |  |  |  |  |
Forfeited, expired or cancelled |  | (168,178) |  |  | 36.78 |  |  |  |  |
Outstanding at end of period |  | 1,839,623 |  | $ | 20.42 |  | 6.88 |  | $ | 26,829
Exercisable at end of period |  | 1,471,669 |  | $ | 21.60 |  | 6.37 |  | $ | 22,569
Weighted Average Assumptions | The fair value of stock options granted during the years ended December 31, 2019 and 2018, was estimated based on the following weighted average assumptions for:
|  | Year Ended December 31, |
|  | 2019 |  | 2018 |
Dividend yield |  | 0 | % | 0 | %
Expected volatility |  | 102 | % | 101 | %
Risk-free interest rate |  | 2.42 | % | 2.72 | %
Expected term |  | 5.62 years |  | 5.60 years |
Stock-based Compensation Expense | The following table shows total stock-based compensation expense for stock options, RSUs and ESPP in the consolidated statements of operations and comprehensive loss (in thousands):
|  | Year Ended December 31,
|  | 2019 |  | 2018
Research and development |  | $ | 204 |  | $ | 369
General and administrative |  |  | 4,744 |  |  | 3,533
Total stock-based compensation expense |  | $ | 4,948 |  | $ | 3,902

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