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Research and Collaboration Arrangements - Roche Agreement - Additional Information (Details) - USD ($) $ in Thousands | 1 Months Ended | 12 Months Ended
--- | --- | ---
Nov. 30, 2017 | Dec. 31, 2021 | Dec. 31, 2020
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] |  |  |
Revenues |  | $ 18,038 | $ 13,612
Roche | 2017 Roche Agreement |  |  |
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] |  |  |
Non-refundable upfront payments received | $ 21,000 |  |
Contingent payment for each nominated product beyond the first three | 1,000 |  |
Maximum contingent payment upon exercise of the option to convert a product | 30,000 |  |
Maximum development milestone payments | 223,000 |  |
Development milestone payments relating to Choroideremia | 86,000 |  |
Maximum sales-based milestone payments | $ 123,000 |  |
Non-refundable development milestone payments |  | 10,000 |
Collaborative arrangement adjustment to transaction price |  | 8,700 | 17,000
Collaborative arrangement adjustment to transaction price due to increase in project scope and expected reimbursable costs |  |  | 7,000
Collaborative arrangement adjustment to transaction price due to variable consideration |  |  | 10,000
Increase in revenue recognized related to performance obligations partially satisfied in the beginning of the current year |  | 11,100 | 4,600
Revenues |  | 17,200 | 12,900
Deferred revenue |  | 0 | 14,300
Accounts receivable |  | 0 | 1,100
Aggregate amount of transaction price allocated to remaining performance obligation |  | $ 0 | $ 25,800

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