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Research and Collaboration Arrangements - Summary of Deferred Revenue (Details) - USD ($) $ in Thousands | Dec. 31, 2021 | Dec. 31, 2020
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Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] |  |
Total deferred revenue | $ 5,090 | $ 19,812
uniQure |  |
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] |  |
Total deferred revenue | 3,661 | 4,396
Roche |  |
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] |  |
Total deferred revenue |  | 14,318
CFF |  |
Collaborative Arrangements And Noncollaborative Arrangement Transactions [Line Items] |  |
Total deferred revenue | $ 1,429 | $ 1,098

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