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Commitments and Contingencies - Additional Information (Details) € in Thousands |  |  | 1 Months Ended | 12 Months Ended |
--- | --- | --- | --- | --- | ---
May 16, 2022 USD ($) | Jan. 14, 2014 | Mar. 30, 2026 USD ($) | Sep. 30, 2024 USD ($) | Sep. 30, 2022 USD ($) | Jan. 31, 2022 USD ($) | Jul. 31, 2019 USD ($) | Dec. 31, 2014 USD ($) LicensedProduct | Sep. 30, 2013 USD ($) | Jul. 31, 2011 USD ($) | Nov. 30, 2009 EUR (€) | Dec. 31, 2025 USD ($) OperatingLease | Dec. 31, 2024 USD ($) | Jan. 31, 2014 USD ($)
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Estimated payments due |  |  |  |  |  |  |  |  |  |  |  | $ 24,346,000 |  |
Operating lease liabilities |  |  |  |  |  |  |  |  |  |  |  | 17,377,000 |  |
Research and development expenses |  |  |  |  |  |  |  |  |  |  |  | 90,300,000 | $ 73,302,000 |
National Institute of Health |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
License issue royalty fee |  |  |  |  |  |  |  |  | $ 100,000 |  |  |  |  |
Sales milestone payments |  |  |  |  |  |  |  |  | 600,000 |  |  |  |  |
Development and regulatory milestone payments |  |  |  |  |  |  |  |  | $ 700,000 |  |  |  |  |
Patent licensing reimbursement fees |  |  |  |  |  |  |  |  |  |  |  | 30,000.00 |  |
Expenses for patent licensing fees |  |  |  |  |  |  |  |  |  |  |  | 40,000.00 | 30,000.00 |
Royalty Bearing Patent License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Expenses |  |  |  |  |  |  |  |  |  |  |  | 0 | 100,000 |
License agreement term |  |  |  |  |  |  |  |  |  |  | 10 years |  |  |
Royalty Bearing Patent License Agreement | IND Filings |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Potential milestone payments | € |  |  |  |  |  |  |  |  |  |  | € 10 |  |  |
Clearside |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Expenses |  |  |  |  |  |  |  |  |  |  |  | 0 | 0 |
Upfront payment |  |  |  |  |  |  | $ 100,000 |  |  |  |  |  |  |
Royalties on net sales |  |  |  |  |  |  | 20.00% |  |  |  |  |  |  |
License agreement expiration period |  |  |  |  |  |  | 10 years |  |  |  |  |  |  |
Clearside | Subsequent Event |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Upfront payment |  |  | $ 1,000,000 |  |  |  |  |  |  |  |  |  |  |
One-time payment |  |  | $ 1,500,000 |  |  |  |  |  |  |  |  |  |  |
Clearside | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Aggregate milestones payment |  |  |  |  |  |  | $ 21,000,000 |  |  |  |  |  |  |
LI-COR, Inc. | 2014 Exclusive Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
One-time upfront license issuance fee |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 100,000
Research and development expenses |  |  |  |  |  |  |  |  |  |  |  | 0 | 0 |
License agreement expiration period |  | 10 years |  |  |  |  |  |  |  |  |  |  |  |
LI-COR, Inc. | 2014 Exclusive Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Aggregate milestone payments |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 200,000
LI-COR, Inc. | 2014 Non- Exclusive Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
License issue fee |  |  |  |  |  |  |  | $ 30,000.00 |  |  |  |  |  |
Milestone payments |  |  |  |  |  |  |  |  |  |  |  | 0 | 0 |
LI-COR, Inc. | 2014 Non- Exclusive Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Payments for development and commercial milestones |  |  |  |  |  |  |  | 400,000 |  |  |  |  |  |
Life Technologies |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Non-refundable, Non- creditable fee |  |  |  |  |  |  |  | $ 300,000 |  |  |  |  |  |
Milestone payments |  |  |  |  |  |  |  |  |  |  |  | 0 | 0 |
Number of licensed products | LicensedProduct |  |  |  |  |  |  |  | 5 |  |  |  |  |  |
Initial license fee |  |  |  |  |  |  |  | $ 100,000 |  |  |  |  |  |
Annual development fee |  |  |  |  |  |  |  | 100,000 |  |  |  |  |  |
Initial license fee for first licensed product |  |  |  |  | $ 400,000 |  |  |  |  |  |  |  |  |
Initial license fee for additional licensed product |  |  |  |  | 500,000 |  |  |  |  |  |  |  |  |
Payment for sublicense |  |  |  |  | 200,000 |  |  |  |  |  |  |  |  |
Payment for use of cell line document package |  |  |  |  | 30,000.00 |  |  |  |  |  |  |  |  |
Change of control fees |  |  |  |  | $ 500,000 |  |  | 200,000 |  |  |  |  |  |
One time perpetual license agreement fee |  |  |  |  |  | $ 50,000.00 |  |  |  |  |  |  |  |
Expense related to agreement including sublicense |  |  |  |  |  |  |  |  |  |  |  | 0 | 500,000 |
Dr. John Schiller | 2014 Non- Exclusive Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Required amendment payment |  |  |  | $ 60,000.00 |  |  |  |  |  |  |  |  |  |
Required amendment payment due period |  |  |  | 30 days |  |  |  |  |  |  |  |  |  |
Dr. John Schiller | Collaboration Research and Development Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Annual development fee |  |  |  |  |  |  |  |  |  | $ 30,000.00 |  |  |  |
Agreement term |  |  |  |  |  |  |  |  |  | 2 years |  |  |  |
First payment due |  |  |  |  |  |  |  |  |  | 30 days |  |  |  |
Subsequent payments due |  |  |  |  |  |  |  |  |  | 30 days |  |  |  |
Research collaboration fees |  |  |  |  |  |  |  |  |  |  |  | 0 | 0 |
Dr. John Schiller | Collaboration Research and Development Agreement | 10th Anniversary |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Required amendment payment |  |  |  | $ 50,000.00 |  |  |  |  |  |  |  |  |  |
Required amendment payment due period |  |  |  | 30 days |  |  |  |  |  |  |  |  |  |
Dr. John Schiller | Collaboration Research and Development Agreement | Ninth Amendment |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Expenses |  |  |  |  |  |  |  |  |  |  |  | $ 50,000.00 | 100,000 |
Rakuten | 2014 Non- Exclusive Agreement |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
License issue fee |  |  |  |  |  |  |  | 30,000.00 |  |  |  |  |  |
Rakuten | 2014 Non- Exclusive Agreement | Maximum |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
License issue fee |  |  |  |  |  |  |  | $ 300,000 |  |  |  |  |  |
Office and Laboratory Facility |  |  |  |  |  |  |  |  |  |  |  |  |  |
Lessee Lease Description [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |
Number of operating lease | OperatingLease |  |  |  |  |  |  |  |  |  |  |  | 1 |  |
Estimated payments due | $ 35,200,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Lease commencement date | Aug. 01, 2022 |  |  |  |  |  |  |  |  |  |  |  |  |
Operating lease, option to extend term | 7 years |  |  |  |  |  |  |  |  |  |  |  |  |
Letters of credit | $ 800,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum reimbursement cost | $ 500,000 |  |  |  |  |  |  |  |  |  |  |  |  |
Incurred of expenses related to expansion |  |  |  |  |  |  |  |  |  |  |  |  | $ 500,000 |

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