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Consolidated Statements of Stockholders' Equity - USD ($) shares in Thousands, $ in Thousands | Common Stock [Member] | Additional Paid-in Capital [Member] | Accumulated Other Comprehensive Income (Loss) [Member] | Accumulated Deficit [Member] | Total
--- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2013 | $ 66 | $ 684,413 | $ 14 | $ (615,313) | $ 69,180
Balance, shares at Dec. 31, 2013 | 66,275 |  |  |  |
Net loss |  |  |  | (66,160) | (66,160)
Other comprehensive loss |  |  | (5) |  | (5)
Issuance of common stock in conjunction with equity plans | $ 2 | 3,966 |  |  | 3,968
Issuance of common stock in conjunction with equity plans, shares | 1,901 |  |  |  |
Issuance of common stock in conjunction with "at-the-market" offering, net of issuance costs | $ 6 | 38,017 |  |  | 38,023
Issuance of common stock in conjunction with "at-the-market" offering, net of issuance costs, shares | 5,751 |  |  |  |
Stock-based compensation expense |  | 9,943 |  |  | 9,943
Balance at Dec. 31, 2014 | $ 74 | 736,339 | 9 | (681,473) | 54,949
Balance, shares at Dec. 31, 2014 | 73,927 |  |  |  |
Net loss |  |  |  | (31,696) | (31,696)
Other comprehensive loss |  |  | (16) |  | (16)
Issuance of common stock in conjunction with equity plans | $ 2 | 7,361 |  |  | 7,363
Issuance of common stock in conjunction with equity plans, shares | 1,981 |  |  |  |
Issuance of common stock in conjunction with "at-the-market" offering, net of issuance costs | $ 4 | 29,096 |  |  | 29,100
Issuance of common stock in conjunction with "at-the-market" offering, net of issuance costs, shares | 4,075 |  |  |  |
Stock-based compensation expense |  | 13,840 |  |  | 13,840
Balance at Dec. 31, 2015 | $ 80 | 786,636 | (7) | (713,169) | $ 73,540
Balance, shares at Dec. 31, 2015 | 79,983 |  |  |  | 79,983
Net loss |  |  |  | (74,375) | $ (74,375)
Other comprehensive loss |  |  | 12 |  | 12
Issuance of common stock in conjunction with equity plans | $ 2 | 7,727 |  |  | 7,729
Issuance of common stock in conjunction with equity plans, shares | 2,004 |  |  |  |
Issuance of common stock in conjunction with "at-the-market" offering, net of issuance costs | $ 7 | 58,193 |  |  | 58,200
Issuance of common stock in conjunction with "at-the-market" offering, net of issuance costs, shares | 6,526 |  |  |  |
Issuance of common stock from exercise of warrant | $ 4 | (4) |  |  |
Issuance of common stock from exercise of warrant, shares | 4,164 |  |  |  |
Stock-based compensation expense |  | 19,562 |  |  | 19,562
Balance at Dec. 31, 2016 | $ 93 | $ 872,114 | $ 5 | $ (787,544) | $ 84,668
Balance, shares at Dec. 31, 2016 | 92,677 |  |  |  | 92,677

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