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Contractual Revenue (Details) - Roche Agreement [Member] - USD ($) | 1 Months Ended | 3 Months Ended | 12 Months Ended |
--- | --- | --- | --- | ---
Dec. 31, 2016 | Sep. 30, 2013 | Dec. 31, 2016 | Sep. 30, 2016 | Jun. 30, 2016 | Mar. 31, 2016 | Dec. 31, 2015 | Sep. 30, 2015 | Jun. 30, 2015 | Mar. 31, 2015 | Dec. 31, 2014 | Sep. 30, 2014 | Jun. 30, 2014 | Mar. 31, 2014 | Dec. 31, 2013 | Dec. 31, 2014 | Jun. 30, 2017
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Non-refundable upfront payment |  | $ 35,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Amortization |  |  |  | $ 3,600,000 | $ 3,600,000 | $ 3,600,000 | $ 3,600,000 | $ 3,600,000 | $ 3,600,000 | $ 3,600,000 | $ 1,700,000 | $ 1,700,000 | $ 1,700,000 | $ 1,700,000 | $ 1,700,000 |  |
Deferred revenue |  |  |  | $ 1,300,000 |  |  |  |  |  |  |  |  |  |  |  |  | $ 0
Collaborative arrangement expiration | Feb. 10, 2017 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Period until expiration upon notice given | 60 days |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, revenue recognized |  |  | $ 1,300,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |
Maximum potential for additional payments to be recognized upon the achievement of certain development milestones |  | $ 40,000,000 |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
First Milestone [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, revenue recognized |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  | $ 10,000,000 |
Second Milestone [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, revenue recognized |  |  |  |  |  |  |  |  | $ 10,000,000 |  |  |  |  |  |  |  |
Third (Final) Milestone [Member] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangements and Non-collaborative Arrangement Transactions [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |  |
Deferred revenue, revenue recognized |  |  |  |  |  |  | $ 20,000,000 |  |  |  |  |  |  |  |  |  |

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