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Consolidated Statements of Stockholders' Equity - USD ($) shares in Thousands, $ in Thousands | At the Market Offering [Member] Common Stock [Member] | At the Market Offering [Member] Additional Paid-in Capital [Member] | At the Market Offering [Member] | Underwritten Public Equity Offering [Member] Common Stock [Member] | Underwritten Public Equity Offering [Member] Additional Paid-in Capital [Member] | Underwritten Public Equity Offering [Member] | Common Stock [Member] | Additional Paid-in Capital [Member] | Accumulated Other Comprehensive Income (Loss) [Member] | Accumulated Deficit [Member] | Total
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Balance at Dec. 31, 2015 |  |  |  |  |  |  | $ 80 | $ 786,636 | $ (7) | $ (713,169) | $ 73,540
Balance, shares at Dec. 31, 2015 |  |  |  |  |  |  | 79,983 |  |  |  |
Net loss |  |  |  |  |  |  |  |  |  | (74,375) | (74,375)
Other comprehensive loss |  |  |  |  |  |  |  |  | 12 |  | 12
Issuance of common stock in conjunction with equity plans |  |  |  |  |  |  | $ 2 | 7,727 |  |  | 7,729
Issuance of common stock in conjunction with equity plans, shares |  |  |  |  |  |  | 2,004 |  |  |  |
Issuance of common stock in conjunction with offering, net of issuance costs | $ 7 | $ 58,193 | $ 58,200 |  |  |  |  |  |  |  |
Issuance of common stock in conjunction with offering, net of issuance costs, shares | 6,526 |  |  |  |  |  |  |  |  |  |
Issuance of common stock from exercise of warrant |  |  |  |  |  |  | $ 4 | (4) |  |  |
Issuance of common stock from exercise of warrant, shares |  |  |  |  |  |  | 4,164 |  |  |  |
Stock-based compensation expense |  |  |  |  |  |  |  | 19,562 |  |  | 19,562
Balance at Dec. 31, 2016 |  |  |  |  |  |  | $ 93 | 872,114 | 5 | (787,544) | 84,668
Balance, shares at Dec. 31, 2016 |  |  |  |  |  |  | 92,677 |  |  |  |
Net loss |  |  |  |  |  |  |  |  |  | (92,189) | (92,189)
Other comprehensive loss |  |  |  |  |  |  |  |  | (37) |  | (37)
Issuance of common stock in conjunction with equity plans |  |  |  |  |  |  | $ 2 | 8,912 |  |  | 8,914
Issuance of common stock in conjunction with equity plans, shares |  |  |  |  |  |  | 2,697 |  |  |  |
Issuance of common stock in conjunction with offering, net of issuance costs | $ 3 | $ 11,862 | $ 11,865 | $ 18 | $ 52,512 | $ 52,530 |  |  |  |  |
Issuance of common stock in conjunction with offering, net of issuance costs, shares | 3,171 |  | 3,200 | 17,732 |  |  |  |  |  |  |
Stock-based compensation expense |  |  |  |  |  |  |  | 20,352 |  |  | 20,352
Balance at Dec. 31, 2017 |  |  |  |  |  |  | $ 116 | 965,752 | (32) | (879,733) | $ 86,103
Balance, shares at Dec. 31, 2017 |  |  |  |  |  |  | 116,277 |  |  |  | 116,277
Net loss |  |  |  |  |  |  |  |  |  | (102,562) | $ (102,562)
Other comprehensive loss |  |  |  |  |  |  |  |  | (4) |  | (4)
ASC606 adoption effect |  |  |  |  |  |  |  |  |  | 189 | 189
Issuance of common stock in conjunction with equity plans |  |  |  |  |  |  | $ 4 | 9,648 |  |  | 9,652
Issuance of common stock in conjunction with equity plans, shares |  |  |  |  |  |  | 3,357 |  |  |  |
Issuance of common stock in conjunction with offering, net of issuance costs |  |  |  | $ 30 | $ 97,500 | $ 97,530 |  |  |  |  |
Issuance of common stock in conjunction with offering, net of issuance costs, shares |  |  |  | 30,610 |  | 30,600 |  |  |  |  |
Stock-based compensation expense |  |  |  |  |  |  |  | 23,153 |  |  | 23,153
Balance at Dec. 31, 2018 |  |  |  |  |  |  | $ 150 | $ 1,096,053 | $ (36) | $ (982,106) | $ 114,061
Balance, shares at Dec. 31, 2018 |  |  |  |  |  |  | 150,244 |  |  |  | 150,244

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