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Consolidated Statements of Stockholders' Equity - USD ($)
shares in Thousands, $ in Thousands
At the Market Offering [Member]
Common Stock
At the Market Offering [Member]
Additional Paid-in Capital [Member]
At the Market Offering [Member]
Underwritten Public Equity Offering [Member]
Common Stock
Underwritten Public Equity Offering [Member]
Additional Paid-in Capital [Member]
Underwritten Public Equity Offering [Member]
Common Stock
Additional Paid-in Capital [Member]
Accumulated Other Comprehensive Income (Loss) [Member]
Accumulated Deficit [Member]
Total
Balance at Dec. 31, 2016             $ 93 $ 872,114 $ 5 $ (787,544) $ 84,668
Balance, shares at Dec. 31, 2016             92,677        
Net loss                   (92,189) (92,189)
Other comprehensive income (loss)                 (37)   (37)
Issuance of common stock in conjunction with equity plans             $ 2 8,912     8,914
Issuance of common stock in conjunction with equity plans, shares             2,697        
Issuance of common stock from offering, net of issuance costs $ 3 $ 11,862 $ 11,865 $ 18 $ 52,512 $ 52,530          
Issuance of common stock from offering, net of issuance costs, shares 3,171   3,200 17,732              
Stock-based compensation expense               20,352     20,352
Balance at Dec. 31, 2017             $ 116 965,752 (32) (879,733) 86,103
Balance, shares at Dec. 31, 2017             116,277        
Net loss                   (102,562) (102,562)
Other comprehensive income (loss)                 (4)   (4)
ASC606 adoption effect                   189 189
Issuance of common stock in conjunction with equity plans             $ 4 9,648     9,652
Issuance of common stock in conjunction with equity plans, shares             3,357        
Issuance of common stock from offering, net of issuance costs       $ 30 $ 97,500 $ 97,530          
Issuance of common stock from offering, net of issuance costs, shares       30,610   30,600          
Stock-based compensation expense               23,153     23,153
Balance at Dec. 31, 2018             $ 150 1,096,053 (36) (982,106) $ 114,061
Balance, shares at Dec. 31, 2018             150,244       150,244
Net loss                   (84,134) $ (84,134)
Other comprehensive income (loss)                 41   41
Issuance of common stock in conjunction with equity plans             $ 3 8,545     8,548
Issuance of common stock in conjunction with equity plans, shares             2,875        
Stock-based compensation expense               16,401     16,401
Balance at Dec. 31, 2019             $ 153 $ 1,120,999 $ 5 $ (1,066,240) $ 54,917
Balance, shares at Dec. 31, 2019             153,119       153,119