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Consolidated Statements of Stockholders' Equity - USD ($) | Common Stock At The Market Offering One | Common Stock At The Market Offering Two | Common Stock | Additional Paid-In Capital At The Market Offering One | Additional Paid-In Capital At The Market Offering Two | Additional Paid-In Capital | Accumulated Deficit Cumulative effect adjustment for adoption of new accounting guidance | Accumulated Deficit | Accumulated Other Comprehensive (Loss) Income | Cumulative effect adjustment for adoption of new accounting guidance | At The Market Offering One | At The Market Offering Two | Total
--- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | --- | ---
Balance, beginning of period at Dec. 31, 2017 |  |  | $ 4,000 |  |  | $ 514,002,000 | $ (474,000) | $ (305,850,000) | $ (76,000) | $ (474,000) |  |  | $ 208,080,000
Balance, beginning of period (in shares) at Dec. 31, 2017 |  |  | 44,507,960 |  |  |  |  |  |  |  |  |  |
Statement of Stockholders' Equity (Deficit) |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for repayment of notes payable |  |  |  |  |  | 22,030,000 |  |  |  |  |  |  | 22,030,000
Issuance of common stock for repayment of notes payable (in shares) |  |  | 636,526 |  |  |  |  |  |  |  |  |  |
Issuance of common stock | $ 1,000 |  |  | $ 48,493,000 | $ 28,387,000 |  |  |  |  |  | $ 48,494,000 | $ 28,387,000 |
Issuance of common stock (in shares) | 1,429,205 | 1,107,000 |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock for asset purchase agreement |  |  |  |  |  | 1,942,000 |  |  |  |  |  |  | 1,942,000
Issuance of common stock for asset purchase agreement (in shares) |  |  | 56,099 |  |  |  |  |  |  |  |  |  |
Exercise of stock options |  |  |  |  |  | 10,328,000 |  |  |  |  |  |  | 10,328,000
Exercise of stock options (in shares) |  |  | 749,294 |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  |  | $ 0 |  |  | 26,598,000 |  | 0 | 0 |  |  |  | 26,598,000
Vesting of restricted common stock awards (in shares) |  |  | 72,000 |  |  |  |  |  |  |  |  |  |
Vesting of employee restricted common stock and common stock subject to repurchase |  |  |  |  |  | 4,000 |  |  | 47,000 |  |  |  | 4,000
Vesting Of Employee Restricted Common Stock And Common Stock Subject To Repurchase (in shares) |  |  | 174,595 |  |  |  |  |  |  |  |  |  |
Purchase of common stock under benefit plans |  |  |  |  |  | 680,000 |  |  |  |  |  |  | 680,000
Purchase of common stock under benefit plans (in shares) |  |  | 26,272 |  |  |  |  |  |  |  |  |  |
Unrealized gain (loss) on marketable debt securities |  |  |  |  |  |  |  |  |  |  |  |  | 47,000
Net loss |  |  |  |  |  |  |  | (109,954,000) |  |  |  |  | (109,954,000)
Balance, end of period at Dec. 31, 2018 |  |  | $ 5,000 |  |  | 652,464,000 | $ 803,000 | (416,278,000) | (29,000) | $ 803,000 |  |  | 236,162,000
Balance, end of period (in shares) at Dec. 31, 2018 |  |  | 48,758,951 |  |  |  |  |  |  |  |  |  |
Statement of Stockholders' Equity (Deficit) |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock |  |  |  | $ 116,356,000 |  |  |  |  |  |  | $ 116,356,000 |  |
Issuance of common stock (in shares) | 4,341,428 |  |  |  |  |  |  |  |  |  |  |  |
Exercise of stock options |  |  |  |  |  | 14,863,000 |  |  |  |  |  |  | 14,863,000
Exercise of stock options (in shares) |  |  | 1,120,186 |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  |  |  |  |  | 27,243,000 |  |  |  |  |  |  | 27,243,000
Vesting of restricted common stock awards (in shares) |  |  | 99,919 |  |  |  |  |  |  |  |  |  |
Purchase of common stock under benefit plans |  |  |  |  |  | 620,000 |  |  |  |  |  |  | 620,000
Purchase of common stock under benefit plans (in shares) |  |  | 35,314 |  |  |  |  |  |  |  |  |  |
Unrealized gain (loss) on marketable debt securities |  |  |  |  |  |  |  |  | 136,000 |  |  |  | 136,000
Net loss |  |  | $ 0 |  |  |  |  | (133,746,000) |  |  |  |  | (133,746,000)
Balance, end of period at Dec. 31, 2019 |  |  | $ 5,000 |  |  | 811,546,000 |  | (549,221,000) | 107,000 |  |  |  | 262,437,000
Balance, end of period (in shares) at Dec. 31, 2019 |  |  | 54,355,798 |  |  |  |  |  |  |  |  |  |
Statement of Stockholders' Equity (Deficit) |  |  |  |  |  |  |  |  |  |  |  |  |
Issuance of common stock |  |  | $ 1,000 |  |  | 203,725,000 |  |  |  |  |  |  | 203,726,000
Issuance of common stock (in shares) |  |  | 6,900,000 |  |  |  |  |  |  |  |  |  |
Exercise of stock options |  |  |  |  |  | 19,500,000 |  |  |  |  |  |  | 19,500,000
Exercise of stock options (in shares) |  |  | 964,412 |  |  |  |  |  |  |  |  |  |
Stock-based compensation expense |  |  |  |  |  | 23,156,000 |  |  |  |  |  |  | 23,156,000
Vesting of restricted common stock awards (in shares) |  |  | 304,638 |  |  |  |  |  |  |  |  |  |
Purchase of common stock under benefit plans |  |  |  |  |  | 896,000 |  |  |  |  |  |  | 896,000
Purchase of common stock under benefit plans (in shares) |  |  | 38,609 |  |  |  |  |  |  |  |  |  |
Unrealized gain (loss) on marketable debt securities |  |  |  |  |  |  |  |  | (153,000) |  |  |  | (153,000)
Net loss |  |  |  |  |  |  |  | (115,976,000) |  |  |  |  | (115,976,000)
Balance, end of period at Dec. 31, 2020 |  |  | $ 6,000 |  |  | $ 1,058,823,000 |  | $ (665,197,000) | $ (46,000) |  |  |  | $ 393,586,000
Balance, end of period (in shares) at Dec. 31, 2020 |  |  | 62,563,457 |  |  |  |  |  |  |  |  |  |

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