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Collaboration and Profit-Sharing Agreements - Beam Therapeutics (Details) - USD ($) $ in Thousands | 1 Months Ended | 3 Months Ended | 12 Months Ended
--- | --- | --- | ---
Oct. 31, 2020 | May 31, 2018 | Dec. 31, 2020 | Sep. 30, 2020 | Jun. 30, 2020 | Mar. 31, 2020 | Dec. 31, 2019 | Sep. 30, 2019 | Jun. 30, 2019 | Mar. 31, 2019 | Dec. 31, 2020 | Dec. 31, 2019 | Dec. 31, 2018
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Realized gain on corporate equity securities | $ 16,400 |  |  |  |  |  |  |  |  |  | $ 16,366 |  |
Collaboration and other research and development revenues |  |  | $ 11,419 | $ 62,841 | $ 10,749 | $ 5,723 | $ 12,284 | $ 3,848 | $ 2,330 | $ 2,069 | $ 90,732 | $ 20,531 | $ 31,937
Revenue type |  |  |  |  |  |  |  |  |  |  | us-gaap:ServiceMember | us-gaap:ServiceMember | us-gaap:ServiceMember
Beam Therapeutics, Inc | License Agreement |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Non-cash consideration aggregate fair value received |  | $ 3,600 |  |  |  |  |  |  |  |  |  |  |
Contract termination notice |  | 90 days |  |  |  |  |  |  |  |  |  |  |
Total transaction price |  |  |  |  |  |  |  |  |  |  |  | $ 3,800 |
Collaboration and other research and development revenues |  |  |  |  |  |  |  |  |  |  | $ 200 | $ 200 |
Revenue type |  |  |  |  |  |  |  |  |  |  | us-gaap:ServiceMember | us-gaap:ServiceMember |
Beam Therapeutics, Inc | License Agreement | Other income (expense) |  |  |  |  |  |  |  |  |  |  |  |  |
Collaborative Arrangement and Arrangement Other than Collaborative [Line Items] |  |  |  |  |  |  |  |  |  |  |  |  |
Realized gain on corporate equity securities |  |  |  |  |  |  |  |  |  |  | $ 16,400 |  |

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