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Selected Quarterly Financial Data (Details) - USD ($)
$ / shares in Units, $ in Thousands
 3 Months Ended 12 Months Ended
Dec. 31, 2020
Sep. 30, 2020
Jun. 30, 2020
Mar. 31, 2020
Dec. 31, 2019
Sep. 30, 2019
Jun. 30, 2019
Mar. 31, 2019
Dec. 31, 2020
Dec. 31, 2019
Dec. 31, 2018
Selected Quarterly Financial Data                      
Total collaboration and other research and development revenues $ 11,419 $ 62,841 $ 10,749 $ 5,723 $ 12,284 $ 3,848 $ 2,330 $ 2,069 $ 90,732 $ 20,531 $ 31,937
Total operating expenses 77,293 53,852 42,088 52,339 51,707 38,436 37,979 33,331 225,572 161,453 145,664
Total other income (expense), net 3,375 (1,170) 7,767 8,892 1,653 1,647 1,863 2,013 18,864 7,176 3,773
Net Loss (62,499) 7,819 (23,572) (37,724) (37,770) (32,941) (33,786) (29,249) (115,976) (133,746) (109,954)
Net (loss) income applicable to common stockholders $ (62,499) $ 7,819 $ (23,572) $ (37,724) $ (37,770) $ (32,941) $ (33,786) $ (29,249) $ (115,976) $ (133,746) $ (109,954)
Net loss per share applicable to common stockholders - basic and diluted         $ (0.74) $ (0.66) $ (0.69) $ (0.60)      
Net (loss) income per share attributable to common shareholders, basic (in dollars per share) $ (1.00) $ 0.13 $ (0.43) $ (0.69)              
Net (loss) income per share attributable to common shareholders, diluted (in dollars per share) $ (1.00) $ 0.12 $ (0.43) $ (0.69)